Muscinae

Haematobosca stimulans, copyright Janet Graham.

Belongs within: Azeliinae.
Contains: Neomyia, MorelliaMesembrina, Musca.

The Muscinae are a group of house flies with a plumose arista, the plumes being more dense (sometimes exclusively) on the upper side (Fan 1992). The lower calypter is broad and posteriorly subtruncate, and the inner corner extends slightly under the scutellum at rest.

The stable flies of the genus Stomoxys and horn flies Haematobia and Haematobosca have the proboscis hardened into a stabbing organ for piercing skin. The palpus is about as long as the proboscis in Haematobia and Haematobosca but considerably shorter in Stomoxys. The arista bears both dorsal and ventral plumes in Haematobosca but dorsal plumes only in Haematobia (Couri 2010).

Non-haematophagous genera of Muscinae include Polietes in which the eyes are densely hairy, and vein M1+2 is almost straight, running parallel to or diverging from R4+5 (Fan 1992). Pyrellia is a metallic-coloured genus in which the face, stem-vein, supra-squamal ridge and infra-alar bulla are all bare (Pont 1973).

Muscinae [Haematobiini, Muscini, Stomoxydinae]KP10
|--+--MorelliaKP10
| `--+--MesembrinaKP10
| `--Polietes Rondani 1866KP10, F92
| |--*P. lardaria (Fabricius 1781) [=Musca lardaria]F92
| |--P. domitor (Harris 1780) [=Musca domitor; incl. M. albolineata Fallén 1823]F92
| |--P. fuscisquamosus Emden 1965 [incl. P. (Pseudomorellia) omeishanensis Fan 1965]F92
| |--P. hirticrura Meade 1887F92
| |--P. koreicus Park & Shinonaga 1985F92
| |--P. nigrolimbata (Bonsdorff 1866) [=Aricia nigrolimbata]F92
| `--P. orientalis Pont 1972F92
`--+--+--Haematobia Lepeletier & Serville in Latreille et al. 1828KP10, C10 [incl. Lyperosia Rondani 1856P73]
| | |--*H. irritans (Linnaeus 1758) [=Conops irritans, *Lyperosia irritans, Stomoxis irritans]P73
| | |--H. exigua de Meijere 1903F92 (see below for synonymy)
| | |--H. minuta (Bezzi 1892) [=Lyperosia minuta]F92
| | |--H. stimulansA71
| | `--H. titillans (Bezzi 1907) [=Lyperosia titillans]F92
| `--+--MuscaKP10
| `--Eudasyphora Townsend 1911KP10, F92
| |--E. cyanella (Meigen 1826) (see below for synonymy)F92
| |--E. cyanicolor (Zetterstedt 1845) [=Pyrellia cyanicolor, Dasyphora cyanicolor]F92
| |--E. dasyprosterna Fan & Qian in Fan 1992F92
| |--E. kempi (Emden 1965) (see below for synonymy)F92
| |--E. pavlovskyi (Zimin 1951)F92
| `--E. tateyamensis Shinonaga 1976F92
`--StomoxyiniC10
|--Haematostoma Malloch 1932F92
|--Haematobosca Bezzi 1907C10
| | i. s.: H. alcisHV87
| |--H. (Haematobosca)F92
| | |--*H. (H.) atripalpis (Bezzi 1895) [=Haematobia atripalpis]F92
| | `--H. (H.) perturbans (Bezzi 1907) [=Siphona perturbans]F92
| |--H. (Bdellolarynx Austen 1909)F92
| | `--H. (B.) sanguinolenta (Austen 1909) (see below for synonymy)F92
| `--H. (Lyperosiops Townsend 1912)F92
| `--H. (L.) stimulans (Meigen 1824) [=Stomoxys stimulans]F92
`--Stomoxys Geoffroy 1762KP10, F92
|--*S. calcitrans (Linnaeus 1758) [=Conops calcitrans]F92
|--S. asiliformisR26
|--S. dorsalisR26
|--S. indicus Picard 1908F92
|--S. nigraP73
| |--S. n. nigraP73
| `--S. n. bilineataP73
|--S. sitiens Rondani 1873 (see below for synonymy)F92
|--S. uruma Shinonaga & Kano 1966F92
`--S. varipesP73

Muscinae incertae sedis:
Mitroplatia Enderlein 1935F92
|--*M. pygmaea Enderlein 1935F92
|--M. albisquamaF92
`--M. nivemaculata Fan, Fang & Yang in Fan 1992F92
Dasyphora Robineau-Desvoidy 1830F92
|--D. pratorum (Meigen 1826)RD77, F92 [=Musca pratorumF92; incl. M. agilis Meigen 1826F92, *Dasyphora agilisF92]
|--D. albofasciata (Macquart in Webb & Berthelot 1839) (see below for synonymy)F92
|--D. apicotaeniata Ni 1982F92
|--D. asiatica Zimin 1947F92
|--D. gansuensis Ni 1982F92
|--D. gussakovskii Zimin 1947F92
|--D. paraversicolor Zimin 1951F92
|--D. quadrisetosa Zimin 1951 [incl. D. huiliensis Ni 1982, D. sinensis Ma 1979]F92
|--D. tianshanensis Ni 1982F92
`--D. trichosterna Zimin 1951F92
Rypellia Malloch 1931F92
|--*R. flavipes Malloch 1931F92
|--R. malaisei (Emden 1965) [=Dasyphora (Rypellia) malaisei]F92
`--R. semilutea (Malloch 1923) (see below for synonymy)F92
Pyrellia Robineau-Desvoidy 1830F92
| i. s.: P. attonita Pont 1973P73
| P. habaheensis Fan & Qian in Fan 1992F92
| P. minuta Zimin 1951F92
| P. rapax (Harris 1870) (see below for synonymy)F92
| P. secunda Zimin 1951F92
| P. serena (Meigen 1826) [=Musca serena]P73
| P. tasmaniae Macquart 1846 (see below for synonymy)P73
|--*P. (Pyrellia) vivida Robineau-Desvoidy 1830F92
`--P. (Dasyphoromima Zimin 1951)P73
|--P. (*D.) pavlovskyi Zimin 1951P73
`--P. (D.) occidentalis Peris & Llorente 1963P73
NeomyiaF92
Curranosia Paterson 1957F92
Stygeromyia Austen 1907F92
`--S. maculosa Austen 1907F92
Myiophaea Enderlein 1935C10 [incl. Gordonia Malloch 1926 non Newton 1892P73]
`--M. spissa (Walker 1858)C10 (see below for synonymy)
Pyrellina marsya (Walker 1849) (see below for synonymy)P73

Dasyphora albofasciata (Macquart in Webb & Berthelot 1839) [=Lucilia albofasciata; incl. D. saltuum Rondani 1862]F92

Eudasyphora cyanella (Meigen 1826) [=Musca cyanella, Dasyphora cyanella; incl. Lucilia lasiophthalma Macquart 1834, *Eudasyphora lasiophthalma]F92

Eudasyphora kempi (Emden 1965) [=Dasyphora cyanicolor kempi, E. cyanicolor kempi; incl. D. qinghaiensis Ni 1982]F92

Haematobia exigua de Meijere 1903F92 [=H. irritans exiguaF04, Lyperosia exiguaB88, Siphona exiguaP73; incl. Haematobia australis Malloch 1932F92, Lyperosia flavohirta Brunetti 1910F92]

Haematobosca (Bdellolarynx) sanguinolenta (Austen 1909) [=Bdellolarynx sanguinolenta; incl. Haematobia aculeata Séguy 1935, Siphona carabao Bohart & Gressitt 1946, Haematobia chinensis Patton 1933, Haematobia rufipes Stein 1918]F92

Myiophaea spissa (Walker 1858)C10 [=Bengalia spissaC10; incl. *Gordonia fulvithorax Malloch 1926P73, *Myiophaea ralumensis Enderlein 1935C10]

Pyrellia rapax (Harris 1870) [=Musca rapax; incl. M. aenea Zetterstedt 1838, Pyrellia ignita Robineau-Desvoidy 1830]F92

Pyrellia tasmaniae Macquart 1846 [incl. P. australensis Curran 1927, P. australiensis (l. c.), P. australis (l. c.) non Macquart 1848, P. egle Bigot 1878, Musca extrema Walker 1852, P. extrema]P73

Pyrellina marsya (Walker 1849) [=Musca marsya; incl. Pyrellia analis Macquart 1855, M. chrysotelus Walker 1852]P73

Rypellia semilutea (Malloch 1923) [=Orthellia semilutea; incl. Pyrellia flavipes Enderlein 1934 non R. flavipes Malloch 1931, R. fulvipes Malloch 1932]F92

Stomoxys sitiens Rondani 1873 [=Stomoxia sitiens; incl. Stomoxys dubitalis Malloch 1932, Stomoxys separabilis Séguy 1935]F92

*Type species of generic name indicated

References

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[C10] Couri, M. S. 2010. Key to the Australasian and Oceanian genera of Muscidae (Diptera). Revista Brasileira de Entomologia 54 (4): 529–544.

[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.

[F04] Ferrar, P. 2004. Australian entomology: isolated, or in touch with the rest of the world? Australian Journal of Entomology 43 (3): 329–333.

[HV87] Huckett, H. C., & J. R. Vockeroth. 1987. Muscidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1115–1131. Research Branch, Agriculture Canada.

[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine’s fly. Systematic Entomology 35: 614–635.

[P73] Pont, A. C. 1973. Studies on Australian Muscidae (Diptera). IV. A revision of the subfamilies Muscinae and Stomoxyinae. Australian Journal of Zoology, Supplementary Series 21: 129–296.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

 

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