Belongs within: Tipulidae.
Contains: Limnophila, Gynoplistia, Hexatoma, Chioneini.
The Limnophilini are a group of crane flies with glabrous eyes and tibial spurs. The North American Dactylolabis vestigipennis has the wings reduced to a short stub, sometimes virtually absent (Alexander & Byers 1981). The Neotropical genus Elephantomyia has an elongate rostrum formed from the frons (Gelhaus 2009).
Characters (from Alexander & Byers 1981, as Hexatomini): Eyes glabrous. Terminal segment of palpus short; nasus absent. Tibial spurs present. Wings, if present, with Sc1 complete, free tip of Sc2 absent, tip of R1+2 present, R4 and R5 separate, CuA straight.
<==Limnophilini [Hexatominae, Hexatomini, Limnophila, Limnophilinae]G09
|--+--HexatomaWT11
| `--ChioneiniWT11
`--DactylolabisWT11
|--*D. montanaWT11
|--D. dilutaA40
|--D. gracilistylus Alexander 1926A40
|--D. mokanica Alexander 1940A40
|--D. pemeticaA40
`--D. (Eudactylolabis) vestigipennisAB81
Limnophilini incertae sedis:
Elephantomyia (Elephantomyia)G09
|--E. (E.) orthorhabdaG09
`--E. (E.) westwoodiG09
AtarbaG09
| i. s.: A. australasiaeCM91 [=Ischnothrix australasiaeCM70]
|--A. (Atarba)G09
| |--A. (A.) bifilosaG09
| `--A. (A.) picticornisG09
`--A. (Ischnothrix)G09
|--A. (I.) patensG09
`--A. (I.) voracisG09
Ulomorpha pilosellaAB81, A16
Paradelphomyia (Oxyrhiza) oaxacensisG09
Shannonomyia (Shannonomyia)G09
|--S. (S.) lentaG09
`--S. (S.) lentoidesG09
Epiphragma (Epiphragma)G09
|--E. (E.) fasciapenne [=E. fascipennis]G09
|--E. (E.) gracilicorneG09
|--E. (E.) imitansG09
`--E. (E.) solatrixG09
Phyllolabis encaustaAB81
PolymeraG09
|--P. (Polymera) rogersianaG09
`--P. (Polymerodes) conjunctaG09
Prolimnophila areolataAB81
AustrolimnophilaG09
|--A. (Austrolimnophila)G09
| |--A. (A.) microstictaG09
| `--A. (A.) toxoneuraG09
`--A. (Limnophilella)G09
|--A. (L.) schunkeanaG09
|--A. (L.) subvictorG09
`--A. (L.) victorG09
PseudolimnophilaG09
|--P. inconcussa (Alexander 1913) [=Limnophila inconcussa]A40
|--P. inornataG09
|--P. (Pseudolimnophila) luteipennisG09
|--P. noveboracensisAB81
`--P. siciliana Krzeminski & Skalski 1983P92
Pilaria reconditaAB81
LimnophilaG09
Lecteria (Lecteria)G09
|--L. (L.) acanthosomaG09
`--L. (L.) armillarisG09
SigmatomeraG09
|--S. (Sigmatomera)G09
| |--S. (S.) occultaG09
| `--S. (S.) seguyiG09
`--S. (Austrolimnobia)G09
|--S. (A.) magnificaG09
`--S. (A.) rufaG09
Ellipteroides (Progonomyia)G09
|--E. (P.) nigrobimboG09
|--E. (P.) quinqueplagiataG09
`--E. (P.) slossonaeG09
NeognophomyiaG09
|--N. adaraG09
`--N. hirsutaG09
Eriopterodes latipleuraG09
GynoplistiaS90
Cerozodia Westwood 1835 [incl. Ozocera]S90
`--C. interrupta Westwood 1835S90
Ctenodia fusca [=Gynoplistia fusca]S90
*Type species of generic name indicated
References
[A16] Alexander, C. P. 1916. New or little-known crane-flies from the United States and Canada: Tipulidae, Diptera, Part 2. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (3): 458–514, pls 16–21.
[A40] Alexander, C. P. 1940. New or little-known Tipulidae from eastern China. Notes d’Entomologie Chinoise 8 (1): 1–28.
[AB81] Alexander, C. P., & G. W. Byers. 1981. Tipulidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 153–190. Research Branch, Agriculture Canada.
[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.
[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).
[G09] Gelhaus, J. K. 2009. Tipulidae (crane flies, tipúlidos). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 193–236. NRC Research Press: Ottawa.
[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.
[S90] Skuse, F. A. A. 1890. Diptera of Australia. Part VII.—The Tipulidae brevipalpi. Proceedings of the Linnean Society of New South Wales, series 2, 4 (3): 757–892, pls 21–24.
[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.