Belongs within: Conjunctae.
Contains: Broscini.
The Broscinae are a group of ground beetles characterised by a pedunculate connection between the pro- and mesothorax, and a single supraorbital seta. Members of the genus Apotomus, which have elongate palps and an unmargined prothorax, are found on mudflats adjacent to rivers (Bouchard 2014). The Melaenitae are small to moderate-sized ground beetles with a remarkably long spermathecal diverticulum in females.
<==Broscinae [Broschidae, Broscitae]
|--+--BrosciniR-JC01
| `--Melaenitae [Melaeninae]R-JC01
| |--Melaenus [Melaenini]R-JC01
| | `--M. pigerR-JC01
| `--CymbionotumB14 [CymbionotiniR-JC01]
| `--C. (Procoscinia)B14
| |--C. (P.) fernandezi Ball & Shpeley 2005B14
| `--C. (P.) negreiB14
`--Apotomus Illiger 1807R-JC01, LM87 [Apotominae, Apotomini, Apotomitae]
|--*A. rufus (Rossi 1790) [=Scarites rufus]LM87
|--A. australis Castelnau 1867LM87
|--A. fuscus Motsch 1858B89b
|--A. hirsutulusR-JC01
|--A. mastersii Macleay 1871LM87
|--A. novaehollandiae Castelnau 1867LM87
|--A. reichardti Erwin 1980B14
`--A. xanthotelus Bates 1874B89a
Broscinae incertae sedis:
Brithysternum Macleay 1873LM87
|--*B. calcaratum Macleay 1873LM87
|--B. macleayi Sloane 1910LM87
`--B. nodosum Sloane 1910LM87
Adotela Castelnau 1867LM87
|--A. carenoides Putzeys 1873LM87
|--A. concolor Castelnau 1867LM87
|--A. esmeralda Castelnau 1867LM87
|--A. frenchi Sloane 1890LM87
|--A. nigerrima Macleay 1873LM87
|--A. striolata Putzeys 1873LM87
`--A. viridis (Macleay 1871) [=Promecoderus viridis]LM87
Parroa Castelnau 1867LM87
|--P. apicalis Sloane 1893LM87
|--P. atronitens (Sloane 1890) [=Adotela atronitens]LM87
|--P. australis (Sloane 1890) [=Adotela australis]LM87
|--P. bicolor Castelnau 1867LM87
|--P. carbonaria Castelnau 1867LM87
|--P. grandis Castelnau 1867LM87
|--P. howittii Castelnau 1867LM87
|--P. laevigata Sloane 1893LM87
|--P. noctis Sloane 1893LM87
`--P. violacea Castelnau 1867LM87
Cerotalis Castelnau 1867LM87
|--C. amabilis Sloane 1890LM87
|--C. brachypleura Sloane 1898LM87
|--C. longipes Sloane 1898LM87
|--C. majuscula (Putzeys 1868) [=Promecoderus majusculus]LM87
|--C. semiviolacea Castelnau 1867LM87
|--C. substriata Castelnau 1867LM87
`--C. versicolor Castelnau 1867LM87
Gnathoxys Westwood 1842LM87
|--G. barbatus Macleay 1864LM87
|--G. cicatricosus Reiche 1842LM87
|--G. crassipes Sloane 1898LM87
|--G. foveatus Macleay 1866LM87
|--G. granularis Westwood 1842 [incl. G. blissii Macleay 1866]LM87
|--G. humeralisM80
|--G. insignitus Macleay 1864LM87
|--G. irregularis Westwood 1842LM87
|--G. macleayi Putzeys 1868LM87
|--G. murrumbidgensis Macleay 1865LM87
|--G. obscurus Reiche 1842LM87
|--G. punctipennis Macleay 1873LM87
|--G. submetallicus Macleay 1864LM87
|--G. sulcicollis Sloane 1910LM87
|--G. tesselatus Macleay 1864LM87
`--G. westwoodi Putzeys 1868LM87
Chylnus Sloane 1920 [=Lychnus Putzeys 1868 non Matheson 1832]LM87
`--*C. ater (Putzeys 1868) [=*Lychnus ater; incl. L. strangulatus Bates 1878, L. striatulus Bates 1878]LM87
Percolestus Sloane 1892LM87
`--*P. blackburni Sloane 1892LM87
Percosoma Schaum 1858LM87
|--*P. carenoides (White 1846) (see below for synonymy)LM87
|--P. concolor Sloane 1892LM87
|--P. montanum (Castelnau 1867) [=Mecodema montanum]LM87
|--P. substriatum Moore 1960LM87
`--P. sulcipenne Bates 1878LM87
*Percosoma carenoides (White 1846) [=Broscus carenoides; incl. B. hopei Sturm 1843 (n. n.), Mecodema percoides Castelnau 1867]LM87
*Type species of generic name indicated
References
[B89a] Bates, H. W. 1889a. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.
[B89b] Bates, H.-W. 1889b. Contributions a la faune Indo-Chinoise. 3e mémoire. Carabidae. Annales de la Société Entomologique de France, 6e série 9: 261–286.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.
[M80] Matthews, E. G. 1980. A Guide to the Genera of Beetles of South Australia. Part 1. Archostemata and Adephaga. South Australian Museum: Australia.
[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.