Actinopteri

Belongs within: Actinopterygii.
Contains: Neopterygii, Palaeonisciformes, Acipenseriformes.

The Actinopteri are a clade of fishes including all modern ray-finned fishes with the exception of the bichirs of the Polypteridae. Members of this clade have the premaxilla confined to a region anterior to the orbit, parasphenoid teeth absent, and proximal radials of the dorsal fin substantially enlarged relative to the middle radials (Giles et al. 2017).

<==ActinopteriXG11
    |--+--NeopterygiiXG11
    |  `--+--PalaeonisciformesGX17
    |     `--AcipenserifomesGX17
    `--+--Brachydegma [Brachydegmidae]GX17
       |    `--B. caelatum Dunkle 1939G93
       `--BirgeriidaeG93
            |--Psilichthys selwyni Hall 1900G93
            `--Birgeria Stensiö 1919GX17, G93
                 `--B. groenlandicaGX17

Actinopteri incertae sedis:
  Willomorichthyidae [Rhabdolepididae]G93
    |--Willomorichthys striatulus Gardiner 1969G93
    |--Rhabdolepis macropterus (Bronn 1829)G93
    |--Tienshaniscus longipterus Lui & Wang 1978G93
    |--Sinoniscus macrolepis Lui & Wang 1978G93
    `--Bendenius Traquair 1878G93
         `--B. deneensis (Traquair 1878)G93
  Phanerorhynchus [Phanerorhynchidae, Phanerorhynchiformes]G93
    `--P. armatus Gill 1923G93
  HoluridaeG93
    |--Holurus parki Traquair 1881G93
    |--Australichthys longidorsalis Gardiner 1969G93
    |--Holuropsis yavorskyi Berg 1947G93
    `--PalaeonisconotusG93
  AtherstoniidaeG93
    |--Aestuarichthys fulcratus Gardiner 1969G93
    `--Atherstonia Woodward 1889G93
         `--A. australis Woodward 1902F71
  Dwykia [Dwykiidae]G93
    `--D. analensis Gardiner 1969G93
  Phanerosteon [Carbovelidae]G93
    `--P. mirabile Traquair 1881G93
  StrepheoschemidaeG93
    |--Strepheoschema fouldenensis White 1927G93
    `--Aetheretmon valentiacum White 1927G93
  Cornuboniscus [Cornuboniscidae]G93
    `--C. budensis White 1939G93
  AmblypteridaeG93
    |--Paramblypterus decorus (Egerton 1850)G93
    |--Amblypterus Agassiz 1833G93
    |--Amblypterina Berg 1940G93
    |--Gardinerichthys tewarii Gupta et al. 1978G93
    `--Korutichthys korutensis Kazantseva 1980G93
  HaplolepididaeG93
    |--Haplolepis Miller 1892G93
    |--Pyritocephalus sculptus Fric 1894G93
    `--Protohaplolepis scoticusC93
  Tarrasius [Tarrasiidae, Tarrasiiformes]G93
    `--T. problematicus Traquair 1881G93
  AeduelliidaeG93
    |--Aeduella blainvillei (Agassiz 1833)G93
    |--Westollia crassus (Pholig 1892)G93
    |--Igornella Heyler 1969G93
    |--Decazella Heyler 1969G93
    |--Burbonella Heyler 1969G93
    `--Palaeothrissum Blainville 1818G93
  Cryphiolepis [Cryphiolepididae]G93
    `--C. striatus (Traquair 1881)G93
  Urosthenes [Urosthenidae]G93
    |--U. australis Dana 1848G93
    `--U. latus Woodward 1931G93
  Lawnia [Lawniidae]G93
    `--L. taylorensis Wilson 1953G93
  TrissolepididaeG93
    |--Sceletophorus biserialis Frič 1894G93
    `--Sphaerolepis kounoviensis Frič 1877G93
  Commentrya [Commentryidae]G93
    `--C. traquairi Sauvage 1888G93

*Type species of generic name indicated

References

[C93] Coates, M. I. 1993. New actinopterygian fish from the Namurian Manse Burn Formation of Bearsden, Scotland. Palaeontology 36: 123–146.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[G93] Gardiner, B. G. 1993. Osteichthyes: basal actinopterygians. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 611–619. Chapman & Hall: London.

[GX17] Giles, S., G.-H. Xu, T. J. Near & M. Friedman. 2017. Early members of ‘living fossil’ lineage imply later origin of modern ray-finned fishes. Nature 549: 265–268.

[XG11] Xu, G.-H. & K.-Q. Gao. 2011. A new scanilepiform from the Lower Triassic of northern Gansu Province, China, and phylogenetic relationships of non-teleostean Actinopterygii. Zoological Journal of the Linnean Society 161: 595–612.