Belongs within: Dytiscidae.
Contains: Ilybius.
The Agabinae is a group of diving beetles sharing the characters of an externally visible scutellum, and metatarsal claws that are equal in length. Members of the Agabini are also united by the presence of a linear series of setae at the anteroventral angle of the metafemur (secondarily lost in some taxa) (Miller 2001). Miller (2001) classified Hydronebrius in the Agabinae, despite the latter’s lack of the aforementioned setal series, due to its similar genitalia to derived members of the Agabini; its relationship to the Agabini requires investigation. The genus Hydrotrupes has also been excluded from Agabinae on the basis of the presence of open mandibular grooves in its larva (in contrast to the closed grooves found in other Agabinae and most other Dytiscidae except Copelatinae) but this separation was not supported by Miller (2001). The type genus Agabus is a primarily Holarctic group of species with a continuous bead along the anterior clypeal margin.
<==Agabinae |--Hydronebrius [Hydronebriini]M01 `--Agabini [Ilybiini]M01 | i. s.: AndonectesM01 | AgametrusM01 | LeuronectesM01 | |--L. curtulusM01 | `--L. gaudichaudiiM01 |--Platynectes Regimbart 1879VB21, NR97 (see below for synonymy) | |--*P. decemnotatus (Aubé 1838) [=Agabus decemnotatus]LM87 | |--P. aenescens Sharp 1882 [incl. P. subaenescens Lea 1899]LM87 | |--P. australicus Guéorguiev 1972LM87 | |--P. bakewelli (Clark 1863) [=Agabus bakewelli; incl. P. obscurus Sharp 1882]LM87 | |--P. beroniS86 | |--P. brownei Guéorguiev 1972 [=P. (*Australonectes) brownei]LM87 | |--P. chapmaniS86 | |--P. darlingtoni Guéorguiev 1972 [incl. P. multimaculatus Watts 1978]LM87 | |--P. decempunctatus (Fabricius 1775)LM87 (see below for synonymy) | |--P. laurianus Watts 1978LM87 | |--‘Colymbetes’ lineatus Redtenbacher in Hügel 1844 [=*Plateocolymbus lineatus]NR97 | |--P. magellanicus (Babington 1841) [=Colymbetes magellanicus, P. (*Neoplatynectes) magellanicus]LM87 | |--P. monostigma (Hope 1842) [=Colymbetes monostigma; incl. Agabus latissimus Clark 1863]LM87 | |--P. octodecimmaculatus (Macleay 1825) [=Colymbetes octodecimmaculatus]LM87 | |--P. reticulosus (Clark 1863) [=Agabus reticulosus; incl. P. limbatus Sharp 1882]LM87 | `--P. tasmaniae (Clark 1863) [=Agabus tasmaniae; incl. P. daemeli Sharp 1882, P. gagatinus Lea 1899]LM87 `--+--Hydrotrupes Sharp 1882VB21, RL01 [Hydrotrupinae] | `--H. palpalisM01 `--+--+--+--IlybiusVB21 | | `--+--‘Agabus’ ambiguusM01 | | |--‘Agabus’ bifariusM01 | | |--‘Agabus’ disintegratusM01 | | `--‘Agabus’ punctulatusM01 | `--+--Agabinus Crotch 1873M01, RL01 | | `--A. glabrellusM01 | `--PlatambusVB21 | |--P. (Platambus) maculatusM01 | `--P. (Anagabus) lineatusM01 `--Agabus Leach 1817VB21, M01 (see below for synonymy) | i. s.: A. clavicornisRL01 | A. heydeniRB02 | ‘Gaurodytes’ marginalisR13 | A. paludosusC01 | A. ragazziiG89 |--A. spinipesM01 `--+--+--+--A. cordatusM01 | | `--A. seriatusM01 | `--+--A. punctatusM01 | |--A. semivittatusM01 | `--A. tristisM01 `--+--A. unguicularisM01 |--+--A. antennatusM01 | `--A. crassipes [=Carrhydrus crassipes]M01 |--+--A. approximatusM01 | |--A. griseipennisM01 | `--A. obliteratusM01 `--+--A. japonicusM01 `--+--A. minnesotensisM01 `--A. regularisM01
Agabus Leach 1817VB21, M01 [incl. Acatodes Thomoson 1859RL01, Agabinectes Guignot 1932RL01, Allogabus Guignot 1951RL01, Allonychus Zaitzev 1905RL01, Apator Semenov 1899RL01, Arctodytes Thomson 1874RL01, Asternus Guignot 1931RL01, Carrhydrus Fall 1923RL01, Colymbinectes Falkenström 1936RL01, Dichodytes Thomson 1886RL01, Dichonectes Guignot 1945RL01, Eriglenus Thomson 1859RL01, Gabinectes Guignot 1931RL01, Gaurodytes Thomson 1859RL01, Heteronychus Seidlitz 1887 non Burmeister 1847RL01, Ilybiosoma Crotch 1873RL01, Mesogabus Guéorguiev 1969RL01, Metronectes Sharp 1882RL01, Nebriogabus Guignot 1936RL01, Necticus Hope 1839 (preoc.)RL01, Neonecticus Guignot 1951RL01, Neoplatynectes Vazirani 1970RL01, Parasternus Guignot 1936RL01, Ranagabus Balfour-Browne 1939RL01, Scytodytes Seidlitz 1887RL01, Xanthodytes Seidlitz 1887RL01]
Platynectes Regimbart 1879VB21, NR97 [incl. Australonectes Guéorguiev 1972LM87, Neoplatynectes Guéorguiev 1972 non Vazirani 1970LM87, Plateocolymbus Gistel 1857 (nom. inv.)NR97]
Platynectes decempunctatus (Fabricius 1775)LM87 [=Dytiscus decempunctatusLM87; incl. Agabus lugubris Blanchard 1853LM87, Platynectes lugubrisM97, A. mastersii Macleay 1871LM87, Lancetes ocularis Lea 1895LM87, Platynectes polygrammus Régimbart 1899LM87, Agabus spilopterus Germar 1848LM87]
*Type species of generic name indicated
References
[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.
[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45–92.
[NR97] Nilsson, A. N., & R. E. Roughley. 1997. The genus- and family-group names of the Dytiscidae—additions and corrections. Beiträge zur Entomologie 47 (2): 359–364.
[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.
[RB02] Ribera, I., R. G. Beutel, M. Balke & A. P. Vogler. 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B—Biological Sciences 269: 2351–2356.
[RL01] Roughley, R. E., & D. J. Larson. 2001. Dytiscidae Leach, 1815. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 156–186. CRC Press: Boca Raton.
[S86] Spangler, P. J. 1986. Insecta: Coleoptera. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 622–631. E. J. Brill/Dr W. Backhuys: Leiden.
[VB21] Vasilikopoulos, A., M. Balke, S. Kukowka, J. M. Pflug, S. Martin, K. Meusemann, L. Hendrich, C. Mayer, D. R. Maddison, O. Niehuis, R. G. Beutel & B. Misof. 2021. Phylogenomic analyses clarify the pattern of evolution of Adephaga (Coleoptera) and highlight phylogenetic artefacts due to model misspecification and excessive data trimming. Systematic Entomology 46: 991–1018.