Belongs within: Hypnales.
Contains: Hygrohypnum, Drepanocladus.
The Amblystegiaceae are a clade of mosses primarily found within damp habitats. As currently circumscribed, the Amblystegiaceae are not yet supported by any morphological synapomorphies; it has only been resolved by molecular analysis combining both chloroplast and nuclear loci. Taxa previously included in the Amblystegiaceae were shown by Vanderpoorten et al (2002) to belong to two phylogenetically distinct clades labelled as Amblystegiaceae and Calliergonaceae; characters previously used to unite them were suggested to be convergent adaptations to aquatic habitats. Vanderpoorten et al. (2002) also indicate that the Amblystegiaceae are paraphyletic to the Fabroniaceae, a group of very small mosses often forming silky mats. Many species of Amblystegiaceae have inflated basal alar cells, but these are generally reduced in species inhabiting less damp habitats (Vanderpoorten et al. 2002).
Amblystegium is a genus of mosses found in damp habitats with short cells in the leaves and without a beak on the capsule lid (Dixon 1924).
<==Amblystegiaceae [Amblystegioideae, Campylioideae, Cratoneuraceae] |--+--Cratoneuron (Sull.) Spruce 1867VH02, S04 [incl. Callialaria Ochyra 1989OZB-O03] | | |--C. curvicaule (Jur.) Roth 1899S04 (see below for synonymy) | | | |--C. c. var. curvicauleD24 | | | `--‘Amblystegium’ c. var. strictumD24 | | `--C. filicinum (Hedw.) Spruce 1867S04 (see below for synonymy) | | |--C. f. var. filicinumVH02 | | `--‘Amblystegium’ f. var. whiteheadii Wheldon 1899D24 | `--Palustriella Ochyra 1989VH02, S04 | |--P. commutata (Hedw.) Ochyra 1989S04 (see below for synonymy) | | |--P. c. var. commutataS04 | | |--P. c. var. fluctuans (Schimp.) Ochyra 1989OZB-O03 (see below for synonymy) | | `--P. c. var. sulcata (Lindb.) Ochyra 1989S04 (see below for synonymy) | |--P. decipiens (De Not.) Ochyra 1989S04 (see below for synonymy) | |--P. falcata (Brid.) Hedenäs 1992S04 (see below for synonymy) | | |--P. f. var. falcataD24 | | `--‘Hypnum falcatum’ var. gracilescens non H. aduncum var. gracilescensD24 | `--P. pluristratosaVHJ03 `--+--+--HygrohypnumVH02 | `--+--Serpoleskea (Limpr.) Loeske 1905VH02, OZB-O03 | | |--S. confervoides (Brid.) Loeske 1905OZB-O03 (see below for synonymy) | | `--S. subtilis (Hedw.) Loeske 1905 (see below for synonymy)OZB-O03 | `--FabroniaceaeVH02 | |--Ischyrodon Müll.Hal. 1875SK02 | | `--I. lepturus (Taylor) Schelpe 1970 (see below for synonymy)SK02 | |--Anacamptodon Brid. 1818VH02, S04 | | |--A. fortunei Mitt. 1864SK02 | | |--A. splachnoides (Fröhl. ex Brid.) Brid. 1818S04 (see below for synonymy) | | `--A. wattsii Broth. 1916SK02 | `--Fabronia Raddi 1808SK02 | |--F. australis Hooker 1819 [incl. F. tayloriana]SK02 | |--F. brachydontia [incl. F. wrightii]SK02 | |--F. brachyphylla Müll.Hal. 1895SK02 | |--F. hampeana Sond. 1844 [incl. F. incana, F. tomentosa]SK02 | |--F. minutaN02 | |--F. scottiae Müll.Hal. 1868SK02 | `--F. secundaN02 `--+--+--Campyliadelphus (Kindb.) Chopra 1975VH02, S04 | | |--C. chrysophyllus (Brid.) Chopra 1975S04 (see below for synonymy) | | | |--C. c. var. chrysophyllusD24 | | | `--‘Hypnum chrysophyllum’ var. erectum Bagnall 1896 [incl. Amblystegium geophilum]D24 | | `--C. elodes (Lindb.) Kanda 1975S04 (see below for synonymy) | | |--C. e. var. elodesD24 | | `--‘Hypnum’ e. var. hamulosum non H. hamulosumD24 | `--+--Leptodictyum (Schimp.) Warnst. 1906VH02, SK02 | | `--L. riparium (Hedw.) Warnst. 1906SK02 (see below for synonymy) | | |--L. r. var. ripariumD24 | | `--‘Hypnum’ r. var. splendens non H. splendensD24 | `--Campylium (Sull.) Mitt. 1869VH02, SK02 | |--*C. stellatum (Hedw.) Jensen 1887OZB-O03, S04 (see below for synonymy) | |--C. polygamum (Schimp.) Lange & Jensen 1887OZB-O03, SK02 (see below for synonymy) | |--C. praegracileJ87 | |--C. protensum (Brid.) Kindb. 1894S04 (see below for synonymy) | `--C. velaxum [incl. Sciaromium bellii]M49 `--+--+--+--DrepanocladusVH02 | | `--Cratoneuropsis (Broth.) Fleischer 1923VH02, SK02 | | `--C. relaxa (Hooker & Wilson) Fleischer ex Broth. 1925 (see below for synonymy)SK02 | `--Pseudocalliergon (Limpr.) Loeske 1907VH02, S04 | |--P. sect. PseudocalliergonOZB-O03 | | |--P. trifarium (Weber & Mohr) Loeske 1907OZB-O03, S04 (see below for synonymy) | | `--P. turgescens (Jensen) Loeske 1907OZB-O03, S04 (see below for synonymy) | `--P. sect. Turgidi (Smirnova) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 | `--P. lycopodioides (Brid.) Hedenäs 1990S04 (see below for synonymy) `--+--Hypnobartlettia fontanaVH02 `--Amblystegium Schimp. in Bruch, Schimp. & Gümbel 1953VH02, S04 (see below for synonymy) | i. s.: A. radicale (Beauv.) Schimp. in Bruch et al. 1853S04 (see below for synonymy) |--*A. serpens (Hedw.) Schimp. in Bruch et al. 1853VH02, S04 (see below for synonymy) | |--A. s. var. serpensD24 | `--A. s. var. angustifoliumD24 `--+--A. varium (Hedw.) Lindb. 1870VH02, S04 (see below for synonymy) `--+--A. humile (Beauv.) Crundw. 1981VH02, S04 (see below for synonymy) `--Hygroamblystegium Loeske 1903 (nom. cons.)S04 |--H. fluviatile (Hedw.) Loeske 1903S04 [=Amblystegium fluviatileVH02, Hypnum fluviatileD24] `--H. tenax (Hedw.) Jenn. 1913S04 (see below for synonymy) Amblystegiaceae incertae sedis: Limbella pachylomaVH02 Gradsteinia andicolaVH02 Campylidium (Kindberg) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--*C. hispidulum (Brid.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--C. creperum (Mitt.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--C. lacerulum (Mitt.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--C. porphyreticum (Müll.Hal.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--C. praegracile (Mitt.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--C. quisqueyanum (Buck) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 |--C. sommerfeltii (Myrin) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 (see below for synonymy) |--C. squarrrosobyssoides (Müll.Hal.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 `--C. trichocladum (Taylor) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 Limprichtia Loeske 1907OZB-O03 Nomina nuda: Amblystegium austropalustre Broth. ex Watts 1899SK02 Amblystegium megachaete Dusén 1903D03 Amblystegium perhumile Müll.Hal. ex Paris 1900SK02 Fabronia baileyana Müll.Hal. ex Paris 1900SK02 Fabronia scottiae var. denticulata Müll.Hal. ex Bailey 1892SK02 Fabronia novae-valesiae Geh. ex Burges 1935SK02 Fabronia obtusoacuminata Müll.Hal. ex Muell. 1880SK02 Fabronia sphaerocarpa Müll.Hal. ex Bailey 1883SK02
Amblystegium Schimp. in Bruch, Schimp. & Gümbel 1953VH02, S04 [incl. Orthotheciella (Müll.Hal.) Ochyra 1998SK02]
Amblystegium humile (Beauv.) Crundw. 1981VH02, S04 [=Leptodictyium humile (Beauv.) Ochyra 1981S04, OZB-O03; incl. A. trichopodium var. curvipesOZB-O03, Leptodictyum kochii var. curvipesOZB-O03, A. trichopodium var. kochiiD24, S04, Leptodictyium kochiiS04, A. riparium var. trichopodiumOZB-O03, Leptodictyium trichopodiumS04]
Amblystegium radicale (Beauv.) Schimp. in Bruch et al. 1853S04 [=Campylium radicaleS04, Hypnum radicaleD24; incl. Amblystegium hygrophilumD24, Campylium hygrophilumOZB-O03, Hypnum hygrophilumOZB-O03, Amblystegium oligorrhizonOZB-O03, A. saxatileS04]
*Amblystegium serpens (Hedw.) Schimp. in Bruch et al. 1853VH02, S04 [=Hypnum serpensD24, Stereodon serpensOZB-O03; incl. Amblystegium cashiiOZB-O03, A. serpens var. depauperatumD24, A. juratzkanum Schimp. 1860S04, OZB-O03, A. serpens var. juratzkanumM08, A. perminimumD24, A. serpens var. rigescensOZB-O03, A. serpens var. rigidusculumOZB-O03, A. serpens var. salinum Carrington 1863OZB-O03, D24, A. xerophilumOZB-O03]
Amblystegium varium (Hedw.) Lindb. 1870VH02, S04 [=Hygroamblystegium variumOZB-O03, Hypnum variumOZB-O03, Leskea variaD24, *Orthotheciella varia (Hedw.) Ochyra 1998OZB-O03, SK02, Pseudoleskea variaOZB-O03, Rigodium variumOZB-O03, Stereodon variumOZB-O03; incl. Amblystegium austrohygrophilum Broth. 1916SK02, A. austroserpens Broth. 1915 (preoc.)SK02, A. dicranadelphus (n. n.)SK02, A. varium var. paludosumOZB-O03]
Anacamptodon splachnoides (Fröhl. ex Brid.) Brid. 1818S04 [=Daltonia splachnoides (Smith) Hooker & Taylor 1818SK02, Fabronia splachnoidesOZB-O03, Neckera splachnoidesOZB-O03, Orthotrichum splachnoidesOZB-O03; incl. D. novae-zelandiaeSK02, D. pusilla Hooker & Wilson 1859SK02]
Campyliadelphus chrysophyllus (Brid.) Chopra 1975S04 [=Amblystegium chrysophyllumD24, Campylium chrysophyllumS04, Hypnum chrysophyllumS04, Stereodon chrysophyllusOZB-O03; incl. H. polymorphum Hedwig 1801 (nom. rej.)OZB-O03, Campylium polymorphumOZB-O03]
Campyliadelphus elodes (Lindb.) Kanda 1975S04 [=Hypnum elodesS04, Amblystegium elodesD24, Campylium elodesS04; incl. Campylium elodes f. falcatumOZB-O03, Hypnum elodes var. falcatum non H. falcatumOZB-O03]
Campylidium (Kindberg) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=C. [unranked] Campylidium Kindberg 1897, Hypnum sect. Campylidium (Kindberg) Paris 1900] OZBO03
Campylidium creperum (Mitt.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Stereodon creperus Mitt. 1859]OZB-O03
*Campylidium hispidulum (Brid.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Hypnum hispidulum Brid. 1812, Campylium hispidulum]OZB-O03
Campylidium lacerulum (Mitt.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Stereodon lacerulus Mitt. 1859]OZB-O03
Campylidium porphyreticum (Müll.Hal.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Campylium porphyreticum Müll.Hal. 1898]OZB-O03
Campylidium praegracile (Mitt.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Ctenidium praegracile Mitt. 1869]OZB-O03
Campylidium quisqueyanum (Buck) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Campylium quisqueyanum Buck 1988]OZB-O03
Campylidium sommerfeltii (Myrin) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003OZB-O03 [=Hypnum sommerfeltii Myrin 1832OZB-O03, Campylium hispidulum var. sommerfeltiiD24, H. hispidulum var. sommerfeltiiD24]
Campylidium squarrrosobyssoides (Müll.Hal.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Campylium squarroso-byssoides Müll.Hal. 1897]OZB-O03
Campylidium trichocladum (Taylor) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Pterogonium trichocladum Taylor 1847]OZB-O03
Campylium polygamum (Schimp.) Lange & Jensen 1887OZB-O03, SK02 [=Amblystegium polygamumD24, Campyliadelphus polygamusSK02, Chrysohypnum polygamumOZB-O03, Drepanocladus polygamus (Schimp.) Hedenäs 1997SK02, Hypnum polygamumD24; incl. Rhynchostegiella campylioides Broth. & Watts 1915SK02, Drepanocladus strictifolius Broth. & Watts 1912SK02; incl. Campylium polygamum var. minusOZB-O03, Hypnum polygamum var. minus nec H. collatum var. minus nec H. cupressiforme var. minusD24, Campylium polygamum var. stagnatumOZB-O03, H. polygamum var. stagnatumD24]
Campylium protensum (Brid.) Kindb. 1894S04 [=Amblystegium protensumD24, C. stellatum var. protensum (Brid.) Bryhn 1893OZB-O03, Hypnum stellatum var. protensumD24]
*Campylium stellatum (Hedw.) Jensen 1887OZB-O03, S04 [=Amblystegium stellatumD24, Campyliadelphus stellatusOZB-O03, Hypnum stellatumD24, Stereodon stellatusOZB-O03]
Cratoneuron curvicaule (Jur.) Roth 1899S04 [=Amblystegium curvicauleS04, *Callialaria curvicaulis (Jur.) Ochyra 1989OZB-O03, Cratoneuron filicinum var. curvicauleS04, Hypnum curvicauleD24]
Cratoneuron filicinum (Hedw.) Spruce 1867S04 [=Amblystegium filicinumSK02, Hypnum filicinumD24; incl. Callialaria filicinum var. atrovirensS04, Cratoneuron filicinum var. atrovirensVH02, Cr. filicinum f. falcatulumOZB-O03, Cr. filicinum f. falcatum non Cr. commutatum var. falcatumOZB-O03, Amblystegium fallaxD24, Cr. filicinum var. fallaxS04, Hypnum fallaxOZB-O03, H. formianumD24, Amblystegium filicinum var. gracilescensD24, Cratoneuron filicinum f. gracilescensOZB-O03, Amblystegium filicinum var. trichodesD24, Cr. filicinum f. trichodesOZB-O03, A. filicinum var. vallisclausaeS04, Hypnum vallisclausaeD24, Cratoneuron filicinum f. xerophilumOZB-O03]
Cratoneuropsis relaxa (Hooker & Wilson) Fleischer ex Broth. 1925 [=Campylium relaxum; incl. Hypnum decussatum, Amblystegium decussatum, Campylium decussatum, Cratoneuropsis decussata, Sciaromium elimbatum, S. forsythii, Campylium molle Broth. 1914, Amblystegium novae-valesiae Broth. & Watts 1912, A. subrelaxum, Campylium subrelaxum, Cratoneuropsis subrelaxa (Broth.) Broth. 1925, Hypnum subrelaxum]SK02
Hygroamblystegium tenax (Hedw.) Jenn. 1913S04 [=Amblystegium tenaxVH02; incl. A. irriguumS04, Hygroamblystegium irriguumOZB-O03, Hypnum irriguumD24, Amblystegium irriguum var. spinifoliumD24, Hygroamblystegium tenax f. spinifoliumOZB-O03]
Ischyrodon lepturus (Taylor) Schelpe 1970 [=Hypnum lepturum, Brachythecium lepturum, Fabronia leptura, Juratzkaea leptura, Stereophyllum lepturum]SK02
Leptodictyum riparium (Hedw.) Warnst. 1906SK02 [=Hypnum ripariumSK02, Amblystegium ripariumSK02; incl. A. riparium f. fontinaloidesOZB-O03, A. leptopterygioides (n. n.)SK02, A. riparium f. longifoliumOZB-O03, Hypnum muelleriSK02, A. muelleriSK02, Leptodictyum muelleriSK02, A. riparium f. subsecundumOZB-O03, A. riparium f. tenuisOZB-O03]
Palustriella commutata (Hedw.) Ochyra 1989S04 [=Hypnum commutatumS04, Cratoneuron commutatumS04; incl. Amblystegium glaucumD24, Cr. glaucumOZB-O03]
Palustriella commutata var. fluctuans (Schimp.) Ochyra 1989OZB-O03 [=Amblystegium falcatum var. fluctuansD24; incl. Cratoneuron commutatum var. irrigatumOZB-O03]
Palustriella commutata var. sulcata (Lindb.) Ochyra 1989S04 [=Hypnum sulcatumS04, Amblystegium glaucum var. sulcatumD24, Cratoneuron commutatum var. sulcatumS04; incl. Hypnum breadalbenseD24, H. sulcatum var. subsulcatumD24]
Palustriella decipiens (De Not.) Ochyra 1989S04 [=Hypnum decipiensS04, Amblystegium decipiensD24, Cratoneuron decipiensS04, Thuidium decipiensD24; incl. Hypnum notarisiiD24]
Palustriella falcata (Brid.) Hedenäs 1992S04 [=Hypnum falcatumS04, Amblystegium falcatumD24, Cratoneuron commutatum var. falcatumS04, Palustriella commutata var. falcataS04; incl. Hypnum irrigatumD24, Cratoneuron commutatum var. virescensS04, H. falcatum var. virescensD24, Palustriella commutata var. virescensS04]
Pseudocalliergon sect. Turgidi (Smirnova) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Drepanocladus sect. Turgidus Smirnova 1962]OZB-O03
Pseudocalliergon lycopodioides (Brid.) Hedenäs 1990S04 [=Hypnum lycopodioidesS04, Amblystegium lycopodioidesD24, Drepanocladus lycopodioidesS04, Scorpidium lycopodioidesS04, Stereodon lycopodioidesOZB-O03]
Pseudocalliergon trifarium (Weber & Mohr) Loeske 1907OZB-O03, S04 [=Hypnum trifariumS04, Acrocladium trifariumS04, Amblystegium trifariumD24, Calliergon trifariumS04, Scorpidium trifariumOZB-O03; incl. C. trifarium f. giganteumOZB-O03]
Pseudocalliergon turgescens (Jensen) Loeske 1907OZB-O03, S04 [=Hypnum turgescensS04, Amblystegium turgescensD24, Calliergon turgescensOZB-O03, Scorpidium scorpioides var. turgescensOZB-O03]
Serpoleskea confervoides (Brid.) Loeske 1905OZB-O03 [=Hypnum confervoidesD24, Amblystegiella confervoidesS04, Amblystegium confervoides (Brid.) Schimp. in Bruch et al. 1853S04, Platydictya confervoidesOZB-O03]
Serpoleskea subtilis (Hedw.) Loeske 1905 [=Amblystegiella subtilis, Amblystegium subtile, Hypnum subtile, Leskea subtilis, Platydictya subtilis; incl. Amblystegium tenuissimum]OZB-O03
*Type species of generic name indicated
References
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