Belongs within: Reptantia.
Contains: Paguroidea, Coenobitoidea, Lepidopinae, Albuneinae, Galatheidae, Porcellanidae.
The Anomala (sometimes known as Anomura, a name that other authors would prefer to be associated with a larger grouping also includes the Thalassinida) include the hermit crabs, stone crabs and half crabs. In general, anomalans resemble true crabs (Brachyura) but have a less reduced abdomen.
The best known anomalans are probably the hermit crabs of the Paguridea that commonly carry gastropod shells or other items to cover their largely uncalcified abdomens (though both of the Paguridea superfamilies Paguroidea and Coenobitoidea include non-‘hermit crab’ members). The Hippoidea, sand crabs or mole crabs, have an ovoid body shape and are adapted for burrowing into sandy beaches. The Galatheoidea, squat lobsters and porcelain crabs, are lobster- or crab-like species in which the uropods form a well-developed tail fan. The grey hairy crab Lomis hirta is a grey, flattened, crab-like animal with a covering of brown hairs found in southern Australia.
Synapomorphies (from Dixon et al. 2003): Basal articles of second antenna fitting into a notch in the carapace; coxosternal joints of thoracopods reversed; cheliped rotated so that dactylus is horizontal; first pleopod of female absent.
<==Anomala [Anomalia, Anomaux, Pagurii] |--Lomis [Lomidae, Lomisidae, Lomisoidea, Lomoidea]DAS03 | `--L. hirta (Lamarck 1818)DAS03 |--PagurideaML01 | | i. s.: Chloenopagurus andersoniM01 | | Trizopagurus strigatusM90 | |--PaguroideaML01 | `--CoenobitoideaML01 |--HippoideaB02 | |--AlbuneidaeB02 | | |--LepidopinaeB02 | | `--AlbuneinaeB02 | |--BlepharipodidaeB02 | | |--Lophomastix Benedict 1904 [=Lophmastix (l. c.), Lophomastrix (l. c.), Lophopmastrix (l. c.)]B02 | | | |--*L. diomedeae Benedict 1904B02 | | | |--L. antiqua Schweitzer & Boyko 2000B02 | | | `--L. japonica (Duruflé 1889) (see below for synonymy)B02 | | `--Blepharipoda Randall 1840 (see below for synonymy)B02 | | |--*B. occidentalis Randall 1840 (see below for synonymy)B02 | | |--B. doelloi Schmitt 1942B02 | | |--B. liberata Shen 1949 [=B. libeata (l. c.)]B02 | | `--B. spinosa (Milne Edwards & Lucas 1841) (see below for synonymy)B02 | `--HippidaeB02 | |--Hippa Fabricius 1787 [incl. Remipes Latreille 1806]B01 | | |--*H. adactyla Fabricius 1787B01 | | |--H. cubensisS52 | | |--H. pacifica (Dana 1852)TSH09 | | |--‘Remipes’ scutellatusF15 | | `--‘Remipes’ testudinariusG20 | |--Emerita Scopoli 1777B02 | | |--*E. emerita (Linnaeus 1767) [=Cancer emeritus, Hippa emerita]B55 | | |--E. analogaCH97 | | |--E. portoricensis Schmitt 1935DAS03 | | |--E. rathbunae Schmitt 1935B02 | | `--E. talpoida [=Hippa talpoida]S52 | `--Mastigocheirus gracilisS52 `--GalatheoideaCA04 | i. s.: Lophoraninella cretacea (Dames 1886)SL03 |--GalatheidaeCA04 |--AeglidaeMD01 | |--Haumuriaegla glaessneri Feldmann 1984BWW93 | `--Aegla Leach 1821H86 | |--A. cavernicola Türkay 1972H86 | `--A. strinatii Türkay 1972H86 |--UroptychusMG-H11 [ChirostylidaeMD01] | |--U. australis (Henderson 1885)MG-H11 | |--U. flindersi Ahyong & Poore 2004MG-H11 | |--U. gracilimanus (Henderson 1885)MG-H11 | `--U. hesperius Ahyong & Poore 2004MG-H11 `--PorcellanidaeMD01
Blepharipoda Randall 1840 [=Blefaripoda (l. c.), Blepharopoda (l. c.), Blephoripoda (l. c.); incl. Abrote Philippi 1857, Albunhippa Milne Edwards & Lucas 1841, Albunhipa (l. c.), Albunhippe (l. c.)]B02
*Blepharipoda occidentalis Randall 1840 [=Blepharopoda (l. c.) occidentalis, Blephoripoda (l. c.) occidentalis]B02
Blepharipoda spinosa (Milne Edwards & Lucas 1841) [=*Albunhippa spinosa, Blephaopoda speciosa (l. c.), Blepharopoda (l. c.) spinosa; incl. Abrote spinimana Philippi 1857, Blepharopoda (l. c.) spinimana]B02
Lophomastix japonica (Duruflé 1889) [=Blepharopoda (l. c. for Blepharipoda) japonica, Blephacopoda (l. c. for Blepharipoda) japonica, Lophmastix (l. c.) japonica, Lophopmastrix (l. c.) japonica; incl. Lophomastrix (l. c.) brevirostris Urita 1934, Lophomastix brevirostris, Lophomastix tchangsii Yü 1935, Blepharopoda (l. c.) fauriana Bouvier 1898, Blepharipoda fauriana]B02
*Type species of generic name indicated
References
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