Belongs within: Ditrysia.
Contains: Simaethistidae, Obtectomera, Cossidae, Sesiidae, Zygaenoidea, Tortricidae, Choreutidae.
The Apoditrysia are a clade of lepidopterans in which the apodemes of abdominal sternum 2 are specialised with short, large bases (Grimaldi & Engel 2005).
<==Apoditrysia [Pyralites, Tortricomorpha, Zygaenides]
|--+--+--ObtectomeraKP19
| | `--+--CossoideaNK11
| | | |--CossidaeKP19
| | | `--Brachodidae [Atychidae, Pseudocossinae]NK11
| | | |--Miscera androgynaNC91
| | | |--SynechodesNC91
| | | |--EuthorybetaNC91
| | | |--HoplophractisEG99
| | | |--Brachodes appendiculataEG99
| | | |--SagalassaEG99
| | | `--NigilgiaNC91
| | | |--N. adjectellaNC91
| | | `--N. browni Kallies 2013JG19
| | `--+--SesiidaeKP19
| | `--ZygaenoideaKP19
| `--+--TortricidaeKP19
| `--Immidae [Immoidea]KP19
| |--Loxotrochis Meyrick 1906Z21
| |--Moca chlorolepis Walsingham 1900 [=Imma chlorolepis, Tortricomorpha chlorolepis]JG19
| |--BursadellaDK99
| |--BirthanaDK99
| | |--B. cleisNC91
| | `--B. saturataDK99
| `--ImmaKP19
| |--I. acosmaNC91
| |--I. albofasciataDK99
| |--I. lyriferaNC91
| |--I. stilbiotaNC91
| |--I. tetrasciaKP19
| `--I. vaticinaNC91
`--+--Urodidae [Urodoidea]KP19
| |--SpiladarchaDK99
| |--WockiaDK99
| `--UrodusKP19
| |--U. decensKP19
| |--U. parvulaGE05
| `--U. venatellaDK99
`--ChoreutidaeKP19
Apoditrysia incertae sedis:
Oligostigma [Hypocampidae]WM66
`--O. orbitalisWM66
Schreckensteiniidae [Schreckensteinioidea]GE05
|--Schreckensteinia festaliellaDK99
`--CorsocasisDK99
Galacticidae [Galacticoidea]GE05
|--GalacticaDK99
|--TanaoctenaDK99
`--HomadaulaDK99
|--H. anisocentraDK99
`--H. myriospilaNC91
Epermeniidae [Epermeniinae, Epermenioidea, Ochromolopinae]GE05
|--Gnathifera eurybiasNC91
|--Paraepermenia santaliellaNC91
|--ThambotrichaDK99
|--OchromolopisNC91
| |--O. cornutiferaNC91
| `--O. eurybiasC74
`--EpermeniaNC91
|--E. chaerophylellaDK99
|--E. eurybiasC70
`--E. exilisNC91
Platypteryx falcatariaG20
Stygia australisR26 [=Aglaope (Stygia) australisG20]
ProdidactidaeNK11
DouglasiidaeNK11
|--KlimeschiaNC91
`--TinagmaNC91 [incl. Douglasia Stainton 1854DR99, P66, Douglassia (l. c.) non Bartsch 1934P66]
|--T. balteolellumDR99
|--T. leucanthesNC91
|--T. obscurofasciellaDR99
`--T. perdicillumDR99
SimaethistidaeNK11
*Type species of generic name indicated
References
[C70] Common, I. F. B. 1970. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 765–866. Melbourne University Press.
[C74] Common, I. F. B. 1974. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 98–107. Melbourne University Press.
[DR99] Davis, D. R., & G. S. Robinson. 1999. The Tineoidea and Gracillarioidea. In: Kristensen, N. P. (ed.). Handbuch der Zoologie vol. 4. Arthropoda: Insecta pt 35. Lepidoptera, Moths and Butterflies vol. 1. Evolution, Systematics, and Biogeography pp. 91–117. Walter de Gruyter.
[DK99] Dugdale, J. S., N. P. Kristensen, G. S. Robinson & M. J. Scoble. 1999. The smaller Microlepidoptera-grade superfamilies. In: Kristensen, N. P. (ed.). Handbuch der Zoologie vol. 4. Arthropoda: Insecta pt 35. Lepidoptera, Moths and Butterflies vol. 1. Evolution, Systematics, and Biogeography pp. 217–232. Walter de Gruyter.
[EG99] Edwards, E. D., P. Gentili, M. Horak, N. P. Kristensen & E. S. Nielsen. 1999. The cossoid/sesioid assemblage. In: Kristensen, N. P. (ed.). Handbuch der Zoologie vol. 4. Arthropoda: Insecta pt 35. Lepidoptera, Moths and Butterflies vol. 1. Evolution, Systematics, and Biogeography pp. 181–197. Walter de Gruyter.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.
[KP19] Kawahara, A. Y., D. Plotkin, M. Espeland, K. Meusemann, E. F. A. Toussaint, A. Donath, F. Gimnich, P. B. Frandsen, A. Zwick, M. dos Reis, J. R. Barber, R. S. Peters, S. Liu, X. Zhou, C. Mayer, L. Podsiadlowski, C. Storer, J. E. Yack, B. Misof & J. W. Breinholt. 2019. Phylogenomics reveals the evolutionary timing and pattern of butterflies and moths. Proceedings of the National Academy of Sciences of the USA 116 (45): 22657–22663.
[NC91] Nielsen, E. S., & I. F. B. Common. 1991. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 817–915. Melbourne University Press: Carlton (Victoria).
[NK11] Nieukerken, E. J. van, L. Kaila, I. J. Kitching, N. P. Kristensen, D. C. Lees, J. Minet, C. Mitter, M. Mutanen, J. C. Regier, T. J. Simonsen, N. Wahlberg, S.-H. Yen, R. Zahiri, D. Adamski, J. Baixeras, D. Bartsch, B. Å. Bengtsson, J. W. Brown, S. R. Bucheli, D. R. Davis, J. De Prins, W. De Prins, M. E. Epstein, P. Gentili-Poole, C. Gielis, P. Hättenschwiler, A. Hausmann, J. D. Holloway, A. Kallies, O. Karsholt, A. Y. Kawahara, S. Koster, M. V. Kozlov, J. D. Lafontaine, G. Lamas, J.-F. Landry, S. Lee, M. Nuss, K.-T. Park, C. Penz, J. Rota, A. Schintlmeister, B. C. Schmidt, J.-C. Sohn, M. A. Solis, G. M. Tarmann, A. D. Warren, S. Weller, R. V. Yakovlev, V. V. Zolotuhin & A. Zwick. 2011. Order Lepidoptera Linnaeus, 1758. In: Zhang, Z.-Q. (ed.) Animal Biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 212–221.
[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[WM66] Wallace, A. R., & F. Moore. 1866. List of lepidopterous insects collected at Takow, Formosa, by Mr. R. Swinhoe. Proceedings of the Zoological Society of London 1866: 355–365.
[Z21] Zilli, A. 2021. Tabwecala robinsoni gen. nov., sp. nov., from Vanuatu and its systematic postion in the ‘Ophiusini-Poaphilini’ clade (Lepidoptera, Erebidae). Nota Lepidopterologica 44: 193–211.