Arthroleptinae

Belongs within: Laurentobatrachia.

The Arthroleptinae are a group of frogs with horizontal pupils found in sub-Saharan Africa (Frost et al. 2006). Males often have hyperelongate third fingers.

<==Arthroleptinae [Arthroleptina, Arthroleptini]FG06
    |--CoracodichusFG06
    |--Cardioglossa Boulenger 1900FG06
    |    |--+--C. gracilisPW11
    |    |  `--C. schioetziPW11
    |    `--+--C. elegansPW11
    |       `--+--C. gratiosaPW11
    |          `--+--+--C. leucomystaxPW11
    |             |  `--C. occidentalisPW11
    |             `--+--C. pulchraPW11
    |                `--+--C. manengoubaPW11
    |                   `--C. oreasPW11
    `--Arthroleptis Smith 1849 [incl. Schoutedenella De Witte 1921]FG06
         |  i. s.: A. batesiFG06
         |         A. crusculum Angel 1950 [=Schoutedenella crusculum]FG06
         |         A. discodactyla (Laurent 1954) [=Schoutedenella discodactyla]FG06
         |         A. hematogaster (Laurent 1954) [=Schoutedenella hematogaster]FG06
         |         A. lameerei de Witte 1921 [=Schoutedenella lameerei]FG06
         |         A. loveridgei de Witte 1933 [=Schoutedenella loveridgei]FG06
         |         A. milletihorsini Angel 1922 [=Schoutedenella milletihorsini]FG06
         |         A. mossoensis (Laurent 1954) [=Schoutedenella mossoensis]FG06
         |         A. nimbaensis Angel 1950 [=Schoutedenella nimbaensis]FG06
         |         A. phrynoides (Laurent 1976) [=Schoutedenella phrynoides]FG06
         |         A. pyrrhoscelis Laurent 1952 [=Schoutedenella pyrrhoscelis]FG06
         |         A. spinalis Boulenger 1919FG06
         |         A. troglodytes Poynton 1963 [=Schoutedenella troglodytes]FG06
         |         A. vercammeni (Laurent 1954) [=Schoutedella vercammeni]FG06
         |         A. xenochirus Boulenger 1905 [=Schoutedenella xenochirus]FG06
         |         A. zimmeri Ahl 1925FG06
         |--+--A. xenodactylus Boulenger 1909PW11, FG06 [=Schoutedenella senodactylaFG06]
         |  `--+--A. schubotzi Nieden 1911PW11, FG06 [=Schoutedenella schubotziFG06]
         |     `--A. xenodactyloides Hewitt 1933PW11, FG06 [=Schoutedenella xenodactyloidesFG06]
         `--+--+--A. taeniatusPW11 [=Schoutedenella taeniataFG06]
            |  `--+--A. aureoliPW11
            |     `--A. sylvaticus (Laurent 1954)PW11, FG06 [=Schoutedenella sylvaticaFG06]
            `--+--+--A. franceiPW11
               |  `--A. wahlbergiiPW11
               `--+--+--A. tanneriPW11
                  |  `--+--A. reicheiPW11
                  |     `--+--A. affinisPW11
                  |        `--A. nikeaePW11
                  `--+--A. stenodactylusPW11
                     `--+--+--A. adelphusPW11
                        |  `--A. poecilonotusPW11
                        `--+--A. krokosuaPW11
                           `--A. variabilisPW11

*Type species of generic name indicated

References

[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.

[PW11] Pyron, R. A., & J. J. Wiens. 2011. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61: 543–583.