Asterales

Toropapa Alseuosmia macrophylla, copyright Roaming Eye.

Belongs within: Campanulidae.
Contains: Campanulaceae, Stylidiaceae, Menyanthaceae, Goodeniaceae, Asteraceae.

The Asterales are a diverse clade of flowering plants including the composite-flowered plants, bellflowers and related taxa, characterised by the possession of notably small seeds (mobot.org). Members include the Alseuosmiaceae, an Australasian family of evergreen shrubs bearing urn- or salver-shaped flowers and fleshy berries.

Characters (mobot.org): Starch generally absent; apotracheal parenchyma absent; leaves spiral; corolla apiculi inflexed; stamens free from corolla; pollen grains often tricellular; orbicules absent; nectary present; style long; ovules many/carpel, integument >7 cells across, endothelium present, hypostase absent; antipodal cells ephemeral; embryo suspensor filamentous, micropylar and chalazal endosperm haustoria present; x = 9; mitochondrial rpl2 gene lost.

<==Asterales [Asteriflorae, Campanulales, Campanulanae]
| i. s.: Donatia Forster & Forster 1776A61 [DonatiaceaeT00]
| |--*D. fascicularisA61
| `--D. novae-zealandiae Hooker 1853A61
|--+--CampanulaceaeK03
| `--RousseaceaeCM16
| |--Roussea simplexCM16
| `--+--Carpodetus Forster & Forster 1776CM16, A61
| | `--C. serratus Forster & Forster 1776A61 [incl. C. dentatusC06, C. forsteriC06]
| `--+--Abrophyllum ornansCM16
| `--Cuttsia viburneaCM16
`--+--Pentaphragma [Pentaphragmataceae]CM16
| |--P. begoniaefoliaRC02
| `--P. ellipticumCM16
`--+--+--StylidiaceaeCM16
| `--+--MenyanthaceaeK03
| `--+--GoodeniaceaeCM16
| |--AsteraceaeK03
| `--Boopis [Calyceraceae]K03
| |--B. anthemoidesK03
| `--B. gramineaK03
`--+--+--Phelline [Phellinaceae]CM16
| | |--P. comosaK03
| | `--P. lucidaCM16
| `--ArgophyllaceaeK03
| |--ArgophyllumT00
| `--Corokia Cunn. 1839CM16, A61
| |--C. buddleoides Cunn. 1839A61
| | |--C. b. var. buddleoidesA61
| | `--C. b. var. linearis Cheesem. 1925A61
| |--C. cotoneaster Raoul 1844A61
| |--C. macrocarpa Kirk 1899A61
| `--C. × virgata Turrill 1912 [C. buddleoides × C. cotoneaster; incl. C. cheesemanii Carse 1913]A61
`--AlseuosmiaceaeK03
|--PeriomphaleT00
|--CrispilobaT00
|--Wittsteinia [incl. Memecylanthus]T00
`--Alseuosmia Cunn. 1839A61
|--A. atriplicifolia Cunningham 1839A61
|--A. banksii Cunningham 1839A61
|--A. ligustrifolia Cunningham 1839 [=A. linariifolia var. ligustrifolia]A61
|--A. linariifolia Cunningham 1839 [incl. A. hookeria Colenso 1846]A61
|--A. macrophylla Cunningham 1839A61
|--A. palaeiformis Cunningham 1839A61
|--A. pusilla Colenso 1885 [incl. A. quercifolia var. glauca Kirk 1899]A61
`--A. quercifolia Cunningham 1839 [incl. A. ilex Cunningham 1839]A61

*Type species of generic name indicated

References

[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

[CM16] Crowl, A. A., N. W. Miles, C. J. Visger, K. Hansen, T. Ayers, R. Haberle & N. Cellinese. 2016. A global perspective on Campanulaceae: biogeographic, genomic, and floral evolution. American Journal of Botany 103 (2): 233–245.

[K03] Kårehed, J. 2003. The family Pennantiaceae and its relationships to Apiales. Botanical Journal of the Linnean Society 141: 1–24.

[RC02] Ridley, H. N., & C. Curtis. 1902. Malay plant names. Journal of the Straits Branch of the Royal Asiatic Society 38: 39–122.

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441–647.

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