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Belongs within: Conjunctae.
Contains: Elaphropus, Pericompsus, Tachys, Philipis, Bembidion, Peryphus.
The Bembidiini are a group of carabid beetles that generally have reduced terminal segments on the labial and maxillary palps, and two unequally dilated segments in the tarsi of males. The majority of bembidiins are surface-dwelling hygrophiles though some are xerophilous and others are endogeous (Ball & Bousquet 2001).
The cave-dwelling Horologion speokoites from West Virginia is exceptional in having the terminal fourth maxillary palpomere as long as the third, which is not pubescent as in other genera (Ball & Bousquet 2001). The Nearctic Anillaspis, Serranillus, Anillinus, Anillodes and Micranillodes are endogeous genera lacking eyes and living deep in leaf litter or in the upper layers of soil (Ball & Bousquet 2001). Members of the genus Micratopus have truncate elytra and a single pair of supraorbital setae whereas other Bembidiini possess two pairs of supraorbital setae and the elytra are not truncate (Bell 1990). Members of the subtribe Tachyina are characterised by an apicolateral notch on the protibiae, and usually an apical recurrent groove on the elytra. In Bembidiina, the apex of the protibia is truncate or rounded (Ball & Bousquet 2001).
The Tachyina are a cosmopolitan group of ground beetles characterised by an apicolateral notch on the protibiae, and usually an apical recurrent groove on the elytra.
<==Bembidiini [Bembidiides, Bembidiinae] |--Gouleta [Xystosomina]R-JC01 | `--G. cayennensisR-JC01 |--TachyinaR-JC01 | |--PericompsusR-JC01 | |--TachysBB01 | `--PhilipisB02 `--BembidiinaR-JC01 |--BembidionLM87 |--PeryphusR-JC01 `--Notaphus Dejean 1821R-JC01, BB01 |--*N. varius (Olivier 1795)LM87 (see below for synonymy) |--N. brullei (Gemminger & Harold 1868) (see below for synonymy)LM87 `--N. convergens [=Austronotaphus convergens]R-JC01 Bembidiini incertae sedis: Lymnastis Motschulsky 1862B08 |--*L. indicus (Motschulsky 1851) [=Lymnaeum indicum]B08 |--L. atricapillusB08 |--L. brooksi Baehr 2008B08 |--L. coomaniB08 |--L. decorseiB55 |--L. inopsB08 |--L. macropsB55 |--L. niloticusB55 |--L. pilosus Bates 1892 [incl. Tachys setiger Sloane 1903]LM87 `--L. pullulusB08 Brachytachys Basilewsky 1953B55 |--B. curtulus Basilewsky 1953B55 `--B. villiersi Basilewsky 1953B55 SphaerotachysB55 |--S. fumigatusB55 `--S. haemorrhoidalisB55 Micratopus Casey 1914 [Micratopini]BB01 `--M. aenescens (LeConte 1848)BB01 Anillinus Casey 1918 [incl. Troglanillus Jeannel 1963]BB01 `--A. valentinei (Jeannel 1963)BB01 Asaphidion des Gozis 1886BB01 |--A. cyanicorneF05 | |--A. c. cyanicorneF05 | `--A. c. atlanticum Ortuño & Toribio 2005F05 |--A. flavipes (Linnaeus 1761) [=Cicindela flavipes]B55 `--A. stierlini (Heyden 1880)VPB11 Mioptachys Bates 1882 [incl. Tachymenis Motschulsky 1862 (preoc.)]BB01 `--M. flavicauda (Say 1823)B14 Tachyta Kirby 1837B90, LM87 | i. s.: T. autumnalis Bates 1882FS90 | T. subvirensB55 `--T. (Tachyta)LM87 |--T. (T.) nanaLM87 | |--T. n. nanaLM87 | `--T. n. inornatus (Say 1823) [incl. *T. picipes Kirby 1837]LM87 `--T. (T.) brunnipennis (Macleay 1871)LM87 [=Bembidium brunnipenneLM87, Tachys brunnipennisS96] Porotachys Netolitzky 1914BB01 `--P. bisulcatus (Nicolai 1822)BB01 Illaphanus Macleay 1865LB91, LM87 |--*I. stephensii Macleay 1865LM87 `--I. macleayi Lea 1906LM87 Phrypeus Casey 1924BB01 `--P. rickseckeri (Hayward 1897)BB01 ElaphropusBB01 Polyderis Motschulsky 1862 [incl. Microtachys Casey 1918]BB01 |--*P. brevicornis (Chaudoir 1846) [=Tachys brevicornis]LM87 |--P. antigua Erwin 1971BB01 |--P. laevis (Say 1823)BB01 `--P. leleupi Basilewsky 1953B55 Paratachys Casey 1918BB01 `--*P. austinicus Casey 1918LM87 Micranillodes Jeannel 1963BB01 `--M. depressus Jeannel 1963BB01 Anillodes Jeannel 1963BB01 Serranillus Barr 1995BB01 `--S. jeanneli Barr 1995BB01 Anillaspis Casey 1918BB01 Horologion Valentine 1932BB01 `--H. speokoites Valentine 1932BB01
Notaphus brullei (Gemminger & Harold 1868) [=Bembicidium brullei, Bembidion (Notaphus) brullei, Bembidion variegatum Brullé 1838 non Say 1823]LM87
*Notaphus varius (Olivier 1795)LM87 [=Carabus variusLM87, Bembidion (*Notaphus) variumLM87, Elaphrus (Bembidion) variusG20]
*Type species of generic name indicated
References
[B02] Baehr, M. 2002. New species and additional records of the tachyine Philipis Erwin (Coleoptera: Carabidae: Bembidiinae). Memoirs of the Queensland Museum 48: 1–7.
[B08] Baehr, M. 2008. A new cavernicolous and eyeless species of the genus Lymnastis from Christmas Island, Australia (Insecta: Coleoptera: Carabidae: Bembidiinae). Records of the Western Australian Museum 24 (2): 199–203.
[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.
[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.
[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.
[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.
[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.
[S96] Sloane, T. G. 1896. On the Australian Bembidiides referable to the genus Tachys, with the description of a new allied genus Pyrrotachys. Proceedings of the Linnean Society of New South Wales 21 (3): 355–377.
[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.