Belongs within: Ichneumonidae.
Contains: Campoplex, Eriborus, Delopia, Diadegma, Casinaria.
The Campoplegini are a group of ichneumon wasps that are solitary endoparasites of holometabolous insect larvae. The majority of species attack Lepidoptera but exceptions include members of the genus Olesicampe that develop on Symphyta (Gauld 1984). Distinctive genera include Xanthocampoplex, species of which are distinguished by their largely yellow coloration (Gauld 1984).
Characters (Gauld 1984): Small to moderately large, fore wing length 2–12 mm; head and body often granulate, weakly polished and usually predominantly black and red or black in colour, rarely polished and brighter coloured. Clypeus rather weakly separated from face by groove, with margin thin or blunt, usually weakly arcuate, less commonly truncate or with a vestige of a median apical tooth; mandible fairly evenly bidentate, often with a ventral flange; male flagellum without tyloids. Sternaulus absent or short; posterior transverse carina of mesosternum usually complete; propodeum usually with some carinae, often with areae superomedia and petiolaris confluent, forming a long, somewhat inverted U- or V-shaped area, sometimes with carinae reduced and/or with a median longitudinal furrow. Apical edge of fore tibia usually without a tooth on outer side; tarsal claws usually pectinate, sometimes simple, never lobate. Fore wing with 3r-m present or absent, when present enclosing a more or less rhombic or obliquely quadrangular areolet which is often petiolate above; hind wing with Rs from slightly longer than to shorter than r-m; distal abscissa of Cu1 usually absent or weak. First tergite of gaster long, usually more or less cylindrical, anteriorly with slender petiole, only slightly broadened posteriorly; petiolar spiracles far behind centre of tergite, with or without a glymma; gaster laterally compressed, at least in the posterior part of females and often very strongly so for its whole length; tergites 2-4 usually with very weak microsculpture, without coarse punctures, grooves or furrow (except for thyridia); ovipositor short to moderately long, when projecting beyond apex of gaster usually up-curved; apex of ovipositor always with a dorsal subapical notch, often without a nodus and with indistinct apical teeth.
<==Campoplegini [Macrini, Porizontini]G84
|--+--DusonaBS23
| | |--*D. stramineipesG84
| | `--D. egregiaHR11
| `--Venturia Schrottky 1902BS23, G84 (see below for synonymy)
| |--*V. argentina Schrottky 1902G84
| |--‘*Balcarcia’ bergi Brèthes 1922G84
| |--V. canescensG84 (see below for synonymy)
| |--‘*Devorgilla’ dilatata Cameron 1907G84
| `--‘*Notamorphota’ timocraticae Blanchard 1947G84
`--+--Charops Holmgren 1859BS23, G84 [incl. Gongropelma Enderlein 1921G84, Zacharops Viereck 1912G84]
| |--‘Ichneumon’ cantator [incl. Campoplex decipiens, *Charops decipiens]G84
| |--C. annulipes [=*Zacharops annulipes]G84
| |--‘Agrypon’ bicolor [incl. *Gongropelma formosanum Enderlein 1921]G84
| `--C. pulchripesG84
`--+--+--CasinariaBS23
| `--+--CymodusaBS23
| | |--C. cruentataMS01
| | |--C. distinctaBS23
| | |--C. elegans Szépligeti in Mocsáry & Szépligeti 1901MS01
| | |--C. leucoceraMS01
| | `--C. pulchricornis Szépligeti in Mocsáry & Szépligeti 1901MS01
| `--Campoletis Foerster 1869 (see below for synonymy)G84
| |--*C. tibiator [=Mesoleptus tibiator]G84
| |--‘Ichneumon’ dilatator [incl. Campoplex declinator, *Sagaritis declinator, *Sagaritopsis declinator]G84
| |--‘Campoplex’ rapax [=*Anilastus rapax]G84
| |--C. sonorensisBS23
| |--C. tasmaniensisG84
| `--‘Campoplex’ viennensis [=*Echphora viennensis, *Ecphoropsis viennensis]G84
`--+--Phobocampe clisiocampaeBS23
`--+--Xanthocampoplex Morley 1913BS23, G84
| |--‘Zachresta’ nigromaculata [incl. *Xanthocampoplex orientalis Morley 1913]G84
| `--X. luteusG84 [incl. X. regis Girault 1925G84, G25]
`--+--Cryptophion inaequalipesBS23
`--+--MicrocharopsBS23
`--Melalophacharops Uchida 1928BS23, G84 [incl. Dichelobosmina Uchida 1928G84]
|--*M. tamanukii Uchida 1928G84
`--‘Charops’ papilionis [incl. *Dichelobosmina tuberculata Uchida 1932]G84
Campoplegini incertae sedis:
ScenocharopsG84
RhachioplexG84
KartikaG84
UrvashiaG84
MenakaG84
LemophagusG84
Porizon harpurusMS01
Eucaphila Gauld 1984G84
`--*E. vulgaris Gauld 1984G84
Neolophron Gauld 1984G84
`--*N. canberrai Gauld 1984G84
Picacharops Gauld 1984G84
`--*P. brevithorax Gauld 1984G84
Slenda Gauld 1984G84
`--*S. ocypeta Gauld 1984G84
Sliochia Gauld 1976G84
`--*S. bala Gauld 1976G84
Nepiera Foerster 1869 [incl. Asinamora Foerster 1869]G84
|--‘Ichneumon’ collector [=*Asinamora collector; incl. Limneria concinna, *Nepiera concinna]G84
`--N. sidnicaG84
Enytus Cameron 1905 [incl. Dioctes Foerster 1869 non Ménétriés 1849, Inareolata Ellinger & Sachtleben 1928]G84
|--*E. maculipes Cameron 1905G84
|--‘Campoplex’ exareolatus [=*Dioctes exareolatus, *Inareolata exareolata]G84
`--‘Bosmina’ rapi [=Dioctes rapi]G33
Sinophorus Foerster 1869 [incl. Campopletidea Viereck 1912, Eulimneria Schmiedeknecht 1907]G84
|--*S. canarsiae [=Limneria canarsiae]G84
|--‘Ichneumon’ albidus [=*Eulimneria albida]G84
`--‘Campoplex’ caradrinae [=*Campopletidea caradrinae]G84
Hyposoter Foerster 1869 (see below for synonymy)G84
|--*H. parorgyiae [=Limnerium parorgyiae]G84
|--H. bombycivorusG84
|--H. didymatorP93
|--‘Banchus’ fugitivus [=*Ameloctonus fugitivus]G84
|--‘*Neozachresta’ havrylenkoi Havrylenko & Winterhalter 1949G84
|--‘*Rhythmonotus’ singularisG84
`--‘*Ischnoscopus’ synchloraeG84
Olesicampe Förster 1869 (see below for synonymy)G84
|--*O. longipes [=Ichneumon longipes, *Limneria longipes, *Limnerium longipes]G84
|--‘Limneria’ albidaMS01
|--O. benefactor Hinz 1969HP-W05
|--‘Limneria’ cothurnata [=*Holocremnus cothurnatus]G84
|--‘Limneria’ fuscicarpaMS01
|--O. geniculata Quednau & Lim 1983HP-W05
|--‘*Omoborus’ kincaidiG84
|--‘*Zaplatystoma’ typicum Viereck 1925G84
`--O. vexata (Holmgren 1860)HP-W05
CampoplexG84
EriborusG84
DelopiaG84
DiadegmaG84
Campoletis Foerster 1869 [incl. Anilasta Thomson 1887, Anilastus Foerster 1869, Echphora Foerster 1869 non Conrad 1843, Ecphoropsis Ashmead 1900, Sagaritis Holmgren 1859 non Billberg 1820, Sagaritopsis Hincks 1944]G84
Hyposoter Foerster 1869 [incl. Ameloctonus Foerster 1869, Ischnoscopus Foerster 1869, Neozachresta Havrylenko & Winterhalter 1949, Rhythmonotus Foerster 1869]G84
Olesicampe Förster 1869 [=Limneria Holmgren 1859 non Adams 1851, Limnerium Ashmead 1900, Olesicampa Thomson 1887; incl. Holocremna Thomson 1887, Holocremnus Foerster 1869, Omoborus Foerster 1869, Zaplatystoma Viereck 1925]G84
Venturia Schrottky 1902BS23, G84 [incl. Balcarcia Brèthes 1922G84, Devorgilla Cameron 1907G84, Exidechthis Walkley 1958G84, Idechthis Foerster 1869 non Hübner 1821G84, Notamorphota Blanchard 1947G84]
Venturia canescensG84 [=Devorgilla canescensA71, Nemeritis canescensA71; incl. V. australicusG84, V. christianaeG84, V. ephestiaG84, V. insularisG84, *Idechthis oahuensisG84, *Exidechthis oahuensisG84]
*Type species of generic name indicated
References
[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.
[BS23] Blaimer, B. B., B. F. Santos, A. Cruaud, M. W. Gates, R. R. Kula, I. Mikó, J.-Y. Rasplus, D. R. Smith, E. J. Talamas, S. G. Brady & M. L. Buffington. 2023. Key innovations and the diversification of Hymenoptera. Nature Communications 14: 1212.
[G84] Gauld, I. D. 1984. An Introduction to the Ichneumonidae of Australia. British Museum (Natural History).
[G25] Girault, A. A. 1925. An essay on when a fly is lovable, the ceremony of baptizing some and unlovely hate. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 187–190).
[G33] Girault, A. A. 1933. Some beauties inhabitant not of commercial boudoirs but of Nature’s bosom, notably new insects. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 299–303).
[HP-W05] Heitland, W., & H. Pschorn-Walcher. 2005. Biology and parasitoids of the peculiar alder sawfly, Platycampus luridiventris (Fallen) (Insecta, Hymenoptera, Tenthredinidae). Senckenbergiana Biologica 85 (2): 215–231.
[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.
[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[P93] Pittaway, A. R. 1993. The Hawkmoths of the Western Palaearctic. Harley Books: Colchester.