Belongs within: Myrtaceae.
Contains: Calytrix, Verticordia.
The Chamelaucieae are a group of mostly smooth-leaved plants bearing dry fruits. Historically, the group has been recognised as including species with indehiscent fruits but molecular analyses support the inclusion of genera with capsular fruits such as Baeckea (Wilson et al. 2005). The type genus, Chamelaucium, is native to south-western Australia; the Geraldton wax C. uncinatum is cultivated as an ornamental. Darwinia species bear flowers in pairs or in compound heads, sometimes surrounded by leafy bracts. Thryptomene species have small leaves in opposite pairs, and white or pink flowers arranged singly or in pairs in leaf axils.
Characters (Wilson et al. 2005): Shrubs, mostly glabrous. Leaves opposite, less commonly spiral. Inflorescences mostly of solitary flowers but sometimes forming corymb-like confloresences; stamens usually few, mostly free but occasionally in clusters opposite the sepals, staminodes present in a few genera; anthers usually adnate, rarely versatile, dehiscing by slits or pores. Ovary inferior 1- to 3-locular; ovules anatropous or hemitropous. Fruit a capsule, or indehiscent; seeds reniform or ovoid to linear; embryo with very small cotyledons on a slender neck appressed to a relatively massive hypocotyl.
<==Chamelaucieae [Baeckeeae, Baeckeineae, Calythriceae, Calytricinae, Chamaelaucieae, Thryptomeneae, Thryptomeninae]WO05
|--HomalocalyxWO05
| |--H. ericaeusLK14
| |--H. polyandrusWO05
| `--H. thryptomenoidesG04a
`--+--Babingtonia tozerensisWO05
|--EuryomyrtusWO05
| |--E. leptospermoidesWO05
| `--E. maideniiG04a
|--ThryptomeneWO05
| |--T. asperaM99
| |--T. baeckeaceaKM08
| |--T. ciliataC08
| |--T. decussataK90
| |--T. kochiiG04b
| |--T. maisonneuviiB05
| |--T. saxicolaWO05
| `--T. urceolarisG04a
`--+--VerticordiaWO05
`--DarwiniaWO05
| i. s.: D. citriodoraJK08
| D. oederoidesGK00
| D. taxifoliaH87
| D. vestitaGK00
|--+--D. capitellataWO05
| `--Actinodium cunninghamiiWO05
`--+--D. fascicularisWO05
`--Homoranthus tropicusWO05
Chamelaucieae incertae sedis:
ChamelauciumWO05
|--C. ciliatumG04b
|--C. oenanthumH18
`--C. uncinatumB88
AlutaWO05
|--A. appressaG04b
`--A. asperaG04a
AstarteaWO05
|--A. ambiguaG04b
`--A. fascicularisGK00
BaeckeaWO05
|--B. camphorosmaeRL05
|--B. crenulataH08
|--B. crispifloraOS04
|--B. cunninghamiiB96
|--B. elderianaG04a
|--B. floribundaM93
|--B. frutescensB14
|--B. gunnianaG67
|--B. muricataM93
|--B. pentagonanthaK90
|--B. preissianaOS04
`--B. pygmaeaGK00
BalaustionWO05
CalytrixWO05
CorynantheraWO05
HypocalymmaWO05
|--H. angustifoliumRL05
|--H. robustumSM06
`--H. strictumGK00
MalleostemonWO05
|--M. pedunculatusK90
`--M. roseusG04a
MicromyrtusWO05
|--M. erichseniiG04b
|--M. peltigeraK90
|--M. racemosaG04b
`--M. triptychaG04b
|--M. t. ssp. triptychaG04b
`--M. t. ssp. elataG04b
OchrospermaWO05
PileanthusWO05
|--P. filifoliusH18
|--P. limacisKM08
|--P. peduncularisS86
`--P. vernicosusH18
Rinzia fumanaWO05, OS04
ScholtziaWO05
|--S. involucrataRL05
|--S. leptanthaK90
|--S. oligandraK90
`--S. umbelliferaK90
TriplarinaWO05
*Type species of generic name indicated
References
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[B05] Beard, J. S. 2005. Drainage evolution in the Lake Disappointment Catchment, Western Australia—a discussion. Journal of the Royal Society of Western Australia 88 (2): 57–64.
[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[C08] Cambage, R. H. 1908. Notes on the native flora of New South Wales. Part VI. Deepwater to Torrington and Emmaville. Proceedings of the Linnean Society of New South Wales 33 (1): 45–65, pls 1–2.
[G04a] Gibson, N. 2004a. Flora and vegetation of the Eastern Goldfields Ranges: part 6. Mt Manning Range. Journal of the Royal Society of Western Australia 87 (2): 35–47.
[G04b] Gibson, N. 2004b. Flora and vegetation of the Eastern Goldfields Ranges: part 7. Middle and South Ironcap, Digger Rock and Hatter Hill. Journal of the Royal Society of Western Australia 87 (2): 49–62.
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[G67] Green, R. H. 1967. The murids and small dasyurids in Tasmania. Parts 1 and 2. Records of the Queen Victoria Museum 28: 1–19.
[H08] Hamilton, A. A. 1908 Notes and exhibits. Proceedings of the Linnean Society of New South Wales 33: 435.
[H87] Haviland, E. 1887. Flowering seasons of Australian plants. No. I—List of plants indigenous in the neighbourhood of Sydney, flowering during July. Proceedings of the Linnean Society of New South Wales, series 2, 1 (4): 1049–1051.
[H18] Houston, T. 2018. A Guide to Native Bees of Australia. CSIRO Publishing.
[JK08] Johnstone, R. E., & T. Kirkby. 2008. Distribution, status, social organisation, movements and conservation of Baudin’s cockatoo (Calyptorhynchus baudinii) in south-west Western Australia. Records of the Western Australian Museum 25 (1): 107–118.
[K90] Keighery, G. J. 1990. Vegetation and flora of Shark Bay, Western Australia. In: Berry, P. F., S. D. Bradshaw & B. R. Wilson (eds) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee pp. 61–87. Western Australian Museum.
[KM08] Keighery, G. J., & W. Muir. 2008. Vegetation and vascular flora of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 11–19.
[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.
[M99] Matthews, M. 1999. Heliothine Moths of Australia: A guide to bollworms and related noctuid groups. CSIRO Publishing.
[M93] Mollemans, F. H. 1993. Drummondita wilsonii, Philotheca langei and P. basistyla (Rutaceae), new species from south-west Western Australia. Nuytsia 9: 95–109.
[OS04] Obbens, F. J., & L. W. Sage. 2004. Vegetation and flora of a diverse upland remnant of the Western Australian wheatbelt (Nature Reserve A21064). Journal of the Royal Society of Western Australia 87 (1): 19–28.
[RL05] Rafferty, C., & B. B. Lamont. 2005. Selective feeding by macropods on vegetation regenerating following fire. Journal of the Royal Society of Western Australia 88 (4): 155–165.
[SM06] Semeniuk, C. A., L. A. Milne, P. Ladd & V. Semeniuk. 2006. Pollen in the surface sediments of wetlands in the Becher Point area, southwestern Australia: a baseline for use in interpreting Holocene sequences. Journal of the Royal Society of Western Australia 89 (1): 27–43.
[S86] Snelling, R. R. 1986. The taxonomy and nomenclature of some Australian paragiine wasps (Hymenoptera: Masaridae). Contributions in Science 378: 1–19.
[WO05] Wilson, P. G., M. M. O’Brien, M. M. Heslewood & C. J. Quinn. 2005. Relationships within Myrtaceae sensu lato based on a matK phylogeny. Plant Systematics and Evolution 251: 3–19.