Belongs within: Culicomorpha.
Contains: Podonomini, Pentaneurini, Procladiini, Macropelopiini, Tanytarsini, Fushunitendipini, Lacusitendipini, Chironomus, Polypedilum, Dicrotendipes, Orthocladiini.
The Chironomidae, commonly known as non-biting midges, are a morphologically diverse family of mostly small, delicate, superficially mosquito-like flies. Larvae of chironomids are most commonly aquatic and the adults are also often found around water. Larvae commonly inhabit silk-lined burrows or tube-like silken shelters that may be fixed to the substrate; some Orthocladiinae and Chironominae construct transportable cases (Spies et al. 2009). Some species are marine, being found in the intertidal zone. Adults of most species have reduced mouthparts, but the relict Southern Hemisphere genus Archaeochlus retains biting mandibles (Colless & McAlpine 1991). Many members of the subfamilies Aphroteniinae, Podonominae and Diamesinae are adapted for living in fast-flowing waters (Oliver 1981).
Among the numerous subfamilies, the Chironominae have wings with crossvein bm-cu absent, vein R2+3 present, fore tarsomere 1 as long as or longer than the fore tibia, and the gonostylus usually rigidly fused to the gonocoxite. The Orthocladiinae have similar venation but fore tarsomere 1 is shorter than the fore tibia, and the gonostylus is flexibly attached to the gonocoxite. Tanypodinae have wings with crossvein bm-cu and usually R2+3 present. R2+3 is absent in Podonominae which have bm-cu close to the middle of the wing (Spies et al. 2009).
Characters (Spies et al. 2009): Adults small to medium-sized, about 1–13 mm long. Ocelli absent; frontal tubercles present or absent. Compound eyes larges, often with dorsomedial extensions but never contiguous. Antennae usually strongly sexually dimorphic; scape a flat ring; pedicel globose (especially enlarged in male); males with 6–15 flagellomeres, females with 4–7 or 10–14; male antennae usually with first and either last or penultimate flagellomere longer than others, flagellum usually densely plumose. Mouthparts not piercing; female mandibles rarely present; proboscis rarely elongate; palpus usually elongate with five (four free) segments. Wing usually narrow, broader in female; wings in resting position held to posterior, either flat or like sloping roof; costa reaching at most to tip of wing; wing margin usually reached by two or three branches of R, one of M and two of Cu. Legs usually elongate with fore legs longest, mid legs shortest; first tarsomeres longer than second; males at rest often hold fore legs aloft like feelers. Postnotum usually with medial longitudinal groove. Abdomen usually much longer than wide, especially slender in males. Larvae elongate, cylindrical (thoracic segments inflated in prepupae); with sclerotised, prognathous, usually non-retractable head capsule, three thoracic and nine abdominal segments, and usually anterior and posterior pair of prolegs bearing spines or claws; antennae various; mouthparts directed anteriorly, with mandibles operating in oblique to horizontal plane, and ventromedially with either transverse, usually toothed plate (hypostoma) or movable, toothed ligula; open spiracles usually absent; terminal segments usually with paired procerci and four anal papillae, sometimes also with short lateral and/or longer ventral tubuli.
<==Chironomidae [Chironomides, Culiciformes, Dixamimidae, Protobibionidae, Rhaetomyiidae, Rhaetomyidae, Tendipedidae]
|--Ulaia [Ulaiinae]BLM02
|--UsambaromyiinaeSA09
|--AenneGE05 [AenneinaeBLM02]
| `--A. triassicaGE05
|--Buchonomyia [Buchonomyiinae]SA09
| |--B. brundiniSA09
| `--B. thienemanniZ02
|--AphroteniinaeO81
| |--AphroteniaGE05
| `--+--AphroteniellaGE05
| `--ParaphroteniaGE05
|--TelmatogetoninaeSA09
| |--Thalassomyia bureniSA09, O81
| |--Paraclunio alaskensisO81
| `--Telmatogeton Schiner 1866S89
| |--*T. sanctipauli Schiner 1868F61
| |--T. alaskensisSA09
| |--T. japonicus Tokunaga 1933 [incl. T. australicus Womersley 1936]F61
| `--T. latipennisSA09
|--PodonominaeSA09
| | i. s.: Oryctochlus vulvanus Kalugina in Kalugina & Kovalev 1985BLM02, RJ93
| | Protobibio jurassicusO81
| |--PodonominiO81
| `--BoreochliniO81
| |--+--Lasiodiamesa brustiGE05, O81
| | `--Trichotanypus posticalisGE05, O81
| `--+--+--BoreochlusGE05
| | `--ParaboreochlusGE05
| `--+--ArchaeochlusGE05
| `--AfrochlusGE05
|--Diamesinae [Prodiamesinae]SA09
| | i. s.: Lobodiamesa campbelliB87
| | Boreoheptagyia luridaO81
| | Pseudodiamesa arcticaO81
| | Pagastia particaO81
| | Hesperodiamesa sequaxO81
| | PotthastiaO81
| | Sympotthastia fulvaO81
| | Odontomesa lutosopraO81
| | Prodiamesa olivaceaO81
| | MonodiamesaO81
| |--Diamesa Meigen 1838S89 [DiamesaeB72, Diamesini]
| |--ProtanypusO81 [ProtanypiniF61]
| | `--P. caudatusO81
| `--HeptagyiniB72
| |--Maoridiamesia harrisiB72, B87
| |--Paraheptagyia tonnoiriB72
| `--Heptagyia Philippi 1865S89
| |--*H. annulipes Philippi 1865F61
| |--H. tasmaniae Freeman 1961F61
| `--H. tonnoiri Freeman 1961F61
|--Tanypodinae [Tanypina]SA09
| | i. s.: Coelopynia Freeman 1961F61
| | `--*C. pruinosa Freeman 1961F61
| |--PentaneuriniSA09
| |--NatarsiaO81 [NatarsiiniSA09]
| |--ProcladiiniSA09
| |--MacropelopiiniSA09
| |--Tanypus Meigen 1803S89 [incl. Pelopia (nom. rej.)O81; TanypodiniSA09]
| | |--T. grodhausi Sublette 1964SA09
| | |--T. maculatusR26
| | |--T. monilisS89
| | |--T. neopunctipennis Sublette 1964SA09
| | `--T. varius [=Chironomus (Tanypus) varius]G20
| `--CoelotanypodiniSA09
| |--Naelotanypus viridisSA09
| |--Clinotanypus Kieffer 1913SA09, F61
| | `--C. crux (Wiedemann 1824) (see below for synonymy)F61
| `--Coelotanypus Kieffer 1913SA09, F61
| |--*C. humeralis (Loew 1866) [=Procladius humeralis]F61
| |--C. africanusF61
| `--C. wirthi Freeman 1961F61
|--Chironominae [Chironomina]SA09
| | i. s.: Spaniotoma Philippi 1865S89
| | `--S. conservataP92
| | Tetraphora Philippi 1865S89
| |--TanytarsiniSA09
| |--FushunitendipiniH02
| |--LacusitendipiniH02
| |--Asiatendipes Hong 2002 [Asiatendipini]H02
| | `--*A. labrosus (Hong 1981) [=Microtendipes labrosus, Eocenitendipes labrosus]H02
| |--PseudochironominiSA09
| | |--Pseudochironomus richardsoniSA09
| | |--AedokritusSA09
| | `--ManoaSA09
| |--LongicopuliniH02
| | |--Longicopula Hong 2002H02
| | | `--*L. xilutianensis Hong 2002H02
| | `--‘Longipedia’ Hong 2002 non Claus 1863H02
| | `--*L. longa Hong 2002H02
| |--FrutexitendipiniH02
| | |--Frutexitendipes Hong 2002H02
| | | `--*F. dongzhouheensis Hong 2002H02
| | |--Noditendipes Hong 2002H02
| | | `--*N. guchengziensis Hong 2002H02
| | `--Dongbeitendipes Hong 2002H02
| | `--*D. foliolatus Hong 2002H02
| |--HamicaudiniH02
| | |--Hamicaudus Hong 2002H02
| | | `--*H. osteneus Hong 2002H02
| | |--Succinaspinus Hong 2002H02
| | | `--*S. orientales (Hong 1981) [=Aspinus orientales, Amberaspinus orientales]H02
| | |--Yangitendipes Hong 2002H02
| | | `--*Y. longivalvatus Hong 2002H02
| | `--Sinaspinus Hong 2002H02
| | |--*S. amblopteres (Hong 1981) [=Aspinus amblopteres, Amberaspinus amblopteres]H02
| | `--S. stenopteres (Hong 1981) [=Aspinus stenopteres, Amberaspinus stenopteres]H02
| `--ChironominiSA09
| |--ChironomusSA09
| |--Harnischia curtilamellataSA09
| |--CyphomellaSA09
| |--AcalcarellaO81
| |--BeckidiaSA09
| |--ChernovskiiaSA09
| |--RobackiaSA09
| |--Demicryptochironomus cuneatusSA09
| |--Cryptotendipes casuariusSA09
| |--Paracladopelma undineSA09
| |--SaetheriaSA09
| |--Paralauterborniella nigrohalterisSA09
| |--Omisus picaSA09
| |--Endochironomus subtendensSA09
| |--GraceusO81
| |--Phaenopsectra coracinaSA09, O81
| |--Pagastiella ostansaSA09, O81
| |--TribelosSA09
| |--Pedionomus beckaeO81
| |--DemeijereaSA09
| |--KloosiaSA09
| |--PelomusSA09
| |--LipiniellaSA09
| |--FissimentumSA09
| |--BeardiusSA09
| |--OukuriellaSA09
| |--Paranilothauma strebulosum [=P. strebulosa]SA09
| |--NeelamiaSA09
| |--StelechomyiaSA09
| |--XestochironomusSA09
| |--EndotribelosSA09
| |--HyporhygmaSA09
| |--Townsia subaberransSA09
| |--PolypedilumSA09
| |--DicrotendipesF61
| |--Imparipecten Freeman 1961F61
| | `--*I. pictipes Freeman 1961F61
| |--Parvitergum Freeman 1961F61
| | `--*P. fuscipenne Freeman 1961F61
| |--Paraborniella Freeman 1961F61
| | `--*P. tonnoiri Freeman 1961F61
| |--ZavreliellaSA09
| | |--Z. longisetaSA09
| | `--Z. marmorataPH10
| |--Paratendipes [Paratendipedidae]SA09
| | |--P. albimanusSA09
| | `--P. laiyangensis Hong & Wang 1990RJ93
| |--GlyptotendipesSA09
| | |--G. barbipesSA09
| | `--G. gripekoveniCS77
| |--ParachironomusSA09
| | |--P. hazelriggiSA09
| | `--P. monochromusO81
| |--EinfeldiaSA09
| | |--E. atitlanensis Sublette & Sasa 1994SA09
| | `--E. paganaSA09
| |--GoeldichironomusSA09
| | |--G. amazonicusSA09
| | `--G. holoprasinusSA09
| |--MicrochironomusSA09 [incl. LeptochironomusO81]
| | |--M. deribaeSA09
| | `--M. nigrovittatusSA09
| |--ApedilumSA09
| | |--A. elachistumSA09
| | `--A. subcinctumSA09
| |--Megacentron Freeman 1961F61
| | |--*M. erebeus (Skuse 1889) [=Chironomus erebeus]F61
| | `--M. cuneicalcar [=Pseudochironomus cuneicalcar]F61
| |--Harrisius Freeman 1959F61
| | |--*H. pallidus Freeman 1959F61
| | `--H. montanus (Skuse 1889) [=Tanytarsus montanus; incl. T. cereolus Skuse 1889]F61
| |--Skusella Freeman 1961F61
| | |--*S. subvittata (Skuse 1889) [=Chironomus subvittatus]F61
| | `--S. pallidipes [=Kribiomimus pallidipes, Lauterborniella pallidipes]F61
| |--Microtendipes Kieffer 1915SA09, H02
| | |--M. chloris [incl. Tendipes abbreviatus, *M. abbreviatus]F61
| | |--M. pedellusSA09
| | `--M. umbrosus Freeman 1955F61
| |--CladopelmaSA09 [incl. CryptocladopelmaO81]
| | |--C. curtivalvaPH10
| | |--C. forcipisSA09
| | `--C. spectabileSA09
| |--AxarusSA09
| | |--A. festivusSA09
| | |--A. rogersiSA09
| | `--A. scopulaSA09
| |--Conochironomus Freeman 1961F61
| | |--*C. acutistilus (Freeman 1955) (see below for synonymy)F61
| | |--C. avicula [=Chironomus (Endochironomus) avicula]F61
| | `--C. cygnusPH10
| |--Stenochironomus Kieffer 1919SA09, F61
| | |--*S. gibbus (Fabricius 1794) [=Tipula gibbus]F61
| | |--S. anomalus Freeman 1961F61
| | |--S. cinctusF61
| | `--S. watsoni Freeman 1961F61
| |--Lauterborniella Bause 1914SA09, F61
| | |--*L. agrayloides (Kieffer 1911) [=Tanytarsus agrayloides]F61
| | |--L. fuscoguttata (Kieffer 1922) [=Polypedilum fuscoguttatum]F61
| | |--L. marmorataF61
| | `--L. varipennisF61
| |--Xenochironomus Kieffer 1921F61
| | |--*X. xenolabis (Kieffer 1916) [=Chironomus xenolabis]F61
| | |--X. australiensis (Freeman 1961) [=Chironomus (Xenochironomus) australiensis]F61
| | |--X. festivusO81
| | `--X. scopulaO81
| |--NilothaumaSA09
| | |--N. aletaSA09
| | |--N. babiyiSA09
| | |--N. brayiR86
| | `--N. duenaSA09
| |--Riethia Kieffer 1917F61
| | |--*R. stictoptera Kieffer 1917 [incl. R. asticta Kieffer 1917]F61
| | |--R. cinctipes Freeman 1961F61
| | |--R. plumosa Freeman 1961F61
| | `--R. zeylandica Freeman 1959F61
| |--Stictochironomus Kieffer 1919SA09, F61
| | |--*S. pictulus (Meigen 1830) [=Chironomus pictulus]F61
| | |--S. fluviaticus (Skuse 1889) [=Chironomus fluviaticus]F61
| | |--S. histrioWP99
| | |--S. illawarra Freeman 1961F61
| | `--S. puripennisF61
| |--Kiefferulus Goetghebuer 1922F61
| | |--*K. tendipediformis (Goetghebuer 1921) [=Tanytarsus tendipediformis]F61
| | |--K. chloronotus [=Chironomus (Kiefferulus) chloronotus]F61
| | |--K. duxO81
| | |--K. intertinctus (Skuse 1889) (see below for synonymy)F61
| | |--K. martini (Freeman 1961) [=Chironomus (Kiefferulus) martini]F61
| | `--K. tumidusPH10
| `--Cryptochironomus Kieffer 1918F61
| |--C. angustus (Freeman 1961) [=Chironomus (Cryptochironomus) angustus]F61
| |--C. curtivalva (Kieffer 1917) (see below for synonymy)F61
| |--C. delinificus (Skuse 1889)F61, S89 [=Chironomus delinificusF61]
| |--C. diceras [=Chironomus (Cryptochironomus) diceras]F61
| |--C. digitatusSA09
| |--C. dycei (Freeman 1961) [=Chironomus (Cryptochironomus) dycei]F61
| |--C. forcipatus (Freeman 1954) [=Chironomus (Cryptochironomus) forcipatus]F61
| |--C. griseidorsum [=Chironomus griseidorsum]F61
| `--C. obreptansR86
`--Orthocladiinae [Clunioninae, Corynoneurinae, Hydrobaeninae]SA09
| i. s.: Naonella Boothroyd 1994B04
| |--N. forsythiB04
| `--N. kimihia Boothroyd 2004B04
| Cricotopus van der Wulp 1874SA09, F61
| |--*C. tibialis (Meigen 1818) [=Chironomus tibialis]F61
| |--C. albitarsusPH10
| |--C. albitibia (Walker 1848) [=Chironomus albitibia]F61
| |--C. annuliventris (Skuse 1889) [=Orthocladius annuliventris]F61
| |--C. cingulatusB87
| |--C. laricomalisSA09
| |--C. trifasciatusR86
| `--C. zealandicusB87
| HeleniellaO81
| |--H. curtistilaO81
| `--H. ornaticollisG09
| Paratrichocladius skirwithensisSA09, G09
| Oreadomyia albertaeO81
| Hydrobaenus Fries 1829O81, S89
| |--H. johannseniO81
| |--H. lugubrisO81
| `--H. spinnatisO81
| ThalassosmittiaO81
| Clunio Haliday 1855SA09, S89
| |--*C. marinus Haliday 1855F61
| |--C. adriaticusR13
| |--C. marshalliO81
| |--C. pacificus Edwards 1926F61
| `--C. takahashiiRD77
| EretmopteraO81
| |--E. browniO81
| `--E. murphyiP94
| Tethymyia aptenaO81
| Corynoneura Winnertz 1846SA09, S89
| |--*C. scutellata Winnertz 1846F61
| |--C. australiensis Freeman 1961F61
| |--C. celeripesRD77
| |--C. cristataF61
| |--C. dewulfiF61
| |--C. ferelobataSA09
| `--C. hirvenojaiSA09
| Thienemanniella Kieffer 1911SA09, F61
| |--*T. clavicornis (Kieffer 1925) [=Corynoneura clavicornis]F61
| `--T. trivittata Goetghebuer 1935 [incl. T. analis Freeman 1953]F61
| Symbiocladius equitansSA09, O81
| Acricotopus nitidellusO81
| DiplocladiusF61
| |--*D. (Diplocladius) cultrigerF61
| `--D. (Stictocladius Edwards 1931)F61
| |--D. (*S.) pulchripennis (Edwards 1931) [=*Stictocladius pulchripennis]F61
| |--D. (S.) multiserialis Freeman 1961F61
| |--D. (S.) uniserialis Freeman 1961F61
| `--D. (S.) victoriensis Freeman 1961F61
| Smittia Holmgren 1869SA09, F61
| |--*S. brevipennis Holmgren 1869F61
| |--S. aterrima (Meigen 1818) (see below for synonymy)F61
| |--S. foliaceaR86
| |--S. retracta Freeman 1961F61
| |--S. vernaM83
| |--S. vetusP92
| `--S. wirthi Freeman 1961F61
| Mesocricotopus thienemanniO81
| Nanocladius (Plectopteracoluthus)SA09
| |--N. (P.) bubrachiatusSA09
| `--‘Plecopteracoluthus’ downsiO81
| ParacricotopusO81
| ParacladiusO81
| Halocladius variabilisO81
| Eurycnemus Van der Wulp 1873O81, S89
| `--E. annuliventrisO81
| ParaphaenocladiusSA09
| |--P. exagitansSA09
| | |--P. e. exagitansSA09
| | `--P. e. longipesSA09
| `--P. pseudirritusR86
| HeterotrissocladiusO81
| PseudorthocladiusO81
| |--P. dumicaudusO81
| `--P. filiformisR86
| HeterotanytarsusO81
| Gymnometriocnemus Goetghebuer 1932O81, F61
| |--*G. subnudus (Edwards 1929) [=Metriocnemus subnudus]F61
| |--G. ancudensis [=Metriocnemus ancudensis]F61
| |--G. longicostalis [=Metriocnemus longicostalis]F61
| |--G. nitidulus (Skuse 1889) [=Metriocnemus nitidulus]F61
| |--G. pectinatus Freeman 1961F61
| `--G. wilsoni Freeman 1961F61
| Cardiocladius Kieffer 1912F61
| |--*C. ceylanicus Kieffer 1912F61
| |--C. australiensis Freeman 1961F61
| `--C. obscurusSA09
| Abiskomyia virgoO81
| Psectrocladius simulansO81
| Limnophyes Eaton 1875SA09, S89
| |--*L. pusillus Eaton 1875F61
| |--L. albibasis Freeman 1961F61
| |--L. borealisO81
| |--L. brachytomusSA09
| |--L. guatemalensisSA09
| |--L. pullulus (Skuse 1889) [=Orthocladius pullulus]F61
| `--L. vestitus (Skuse 1889) [=Camptocladius vestitus]F61
| ParachaetocladiusO81
| Synorthocladius semivirensSA09, O81
| TokunagaiaO81
| Clinocladius subparallelusO81
| Chaetocladius Kieffer 1911O81, F61
| |--C. pigerR86
| `--C. stamfordiO81
| Platycladius pleuralisO81
| Baeoctenus bicolorO81
| Psilometriocnemus triannulatusO81
| Oliveria tricornisO81
| ZalutschiaO81
| BryophaenocladiusSA09
| |--B. carusSA09
| |--B. flavoscutellatusSA09
| `--B. (Odontocladius) nigrusR86
| Chasmatonotus Loew 1864O81, S89
| `--C. unimaculatusO81
| KrenosmittiaO81
| Pseudosmittia Goetghebuer 1932SA09, F61
| |--*P. angusta (Edwards 1929) [=Spaniotoma (Smittia) angusta]F61
| |--P. conjuncta [=Smittia conjuncta]F61
| |--P. crassipennis (Skuse 1889) [=Camptocladius crassipennis]F61
| |--P. forcipataSA09
| |--P. imperfecta (Skuse 1889) [=Ceratopogon imperfectus, Spaniotoma (Smittia) imperfectus]F61
| |--P. latifurca Freeman 1961F61
| |--P. paraconjuncta [=Smittia paraconjuncta]F61
| `--P. setavenaO81
| Phycoidella dentolatensO81
| Parakiefferiella variegatusSA09, PH10
| Epoicocladius flavensO81
| Mariocladius subaterrimusO81
| Tempisquitoneura merrillorumSA09
| Onconeura semifimbriataSA09
| OliveiriellaSA09
| Lopescladius (Lopescladius) verruculosusSA09
| Irisobrillia longicosta Oliver 1985SA09
| CompterosmittiaSA09
| |--C. aberransSA09
| `--C. neriusSA09
| ParalimnophyesSA09
| Antillocladius pluspilalusSA09
| Mesosmittia toraSA09
| Lipurometriocnemus glabalusSA09
| DiplosmittiaSA09
| |--D. beluinaSA09
| |--D. carinataSA09
| `--D. forficataSA09
| Belgica antarcticaRD77, N91
| Halirytus Eaton 1875RD77, S89
| Doloplastus Skuse 1889F61, S89
| `--*D. monticola Skuse 1889S89
| Rhinocladius Edwards 1931F61
| |--*R. culicinus Edwards 1929F61
| `--R. tonnoiri Freeman 1961F61
| Allometriocnemus Freeman 1961F61
| |--*A. coloensis Freeman 1961F61
| `--A. pictus Freeman 1961F61
| Nasuticladius Freeman 1961F61
| |--*N. tonnoiri Freeman 1961F61
| |--N. ater Freeman 1961F61
| |--N. niger Freeman 1961F61
| |--N. wilsoni Freeman 1961F61
| `--N. wirthi Freeman 1961F61
| Austrobrillia Freeman 1961F61
| `--*A. longipes Freeman 1961 [=A. nudipennis Freeman 1961 (n. n.)]F61
| Austrocladius Freeman 1961F61
| |--*A. terjugus (Skuse 1889) [=Camptocladius terjugus]F61
| |--A. hamulatus (Edwards 1931) [=Spaniotoma (Orthocladius) hamulata]F61
| |--A. harrisi (Freeman 1959) [=Chaetocladius harrisi]F61
| |--A. heterogeneus (Edwards 1931) [=Spaniotoma (Orthocladius) heterogenea]F61
| |--A. hirtinervis (Edwards 1931) [=Spaniotoma (Orthocladius) hirtinervis]F61
| |--A. numerosus (Skuse 1889) [=Orthocladius numerosus]F61
| |--A. obliquus (Edwards 1931) [=Spaniotoma (Orthocladius) obliqua]F61
| `--A. trichiatus Freeman 1961F61
| Kiefferophyes Freeman 1961F61
| |--*K. invenustulus (Skuse 1889) [=Camptocladius invenustulus, C. investulus (l. c.)]F61
| |--K. leei Freeman 1961F61
| `--K. lobifer Freeman 1961F61
| Camptocladius Van der Wulp 1874F61
| `--*C. stercorarius (De Geer 1776) [=Tipula stercoraria, Spaniotoma (Smittia) stercoraria]F61
| Orthosmittia Goetghebuer 1943F61
| |--*O. brevifurcata (Edwards 1926) [=Orthocladius (Dactylocladius) brevifurcatus]F61
| |--O. bifurcataF61
| |--O. hirtellaF61
| |--O. macrobrachiusF61
| |--O. melanostolaF61
| `--O. reyei Freeman 1961F61
|--OrthocladiiniG09
`--MetriocneminiG09
|--Parametriocnemus Goetghebuer 1932G09, F61
| |--*P. stylatus (Kieffer 1924) [=Metriocnemus stylatus]F61
| |--P. lundbeckiiSA09
| `--P. ornaticornis (Kieffer 1917) [=Metriocnemus ornaticornis]F61
`--Metriocnemus van der Wulp 1874F61
|--*M. albolineatus [=Chironomus albolineatus]F61
|--M. cretatusP92
|--M. fuscipesF61
`--M. hornsbyensis Freeman 1961F61
Chironomidae incertae sedis:
AustrochlusGE05
Euryhapsis subviridisR86
StichtotanytarsusWP99
EopodonomusO81
PachyneuronymphaO81
NimboceraSA09
AllotrissocladiusCM91
Eocenitendipes Evenhuis 1994H02
DolichopelmaH02
CricotopiellaP92
EuhycnemusP92
PalaeotanypusP92
SendeliaP92
JentzschiellaP92
Cretodiamesa taimyricaP92
Electrotenia brundiniP92
Libanochlites neocomicus Brundin 1976P92
Prionomyia femorata [=Ceratopogon femoratus]S89
Labidomyia bipunctata [=Ceratopogon bipunctatus]S89
‘Burmeisteria’ Weyenberg 1886 non Gray 1865S89
Trichocladius Kieffer 1906 (n. d.)F61
Paucispinigera approximataF61
Kribiomyia Kieffer 1921F61
Clinotanypus crux (Wiedemann 1824) [=Tanypus crux; incl. C. australiensis Kieffer 1926, T. ornatus Doleschall 1857, T. pardalis Doleschall 1856]F61
*Conochironomus acutistilus (Freeman 1955) [=Endochironomus acutistilus, Chironomus (Endochironomus) acutistilus]F61
Cryptochironomus curtivalva (Kieffer 1917) [=Chironomus curtivalva; incl. Ch. pulcher Skuse 1889 non Wiedemann 1830]F61
Kiefferulus intertinctus (Skuse 1889) [=Chironomus intertinctus; incl. C. hexhamensis Skuse 1889, C. lativalva Kieffer 1917]F61
Smittia aterrima (Meigen 1818) [=Chironomus aterrimus; incl. S. atra Freeman 1954, Camptocladius macleayi Skuse 1889]F61
*Type species of generic name indicated
References
[BLM02] Blagoderov, V. A., E. D. Lukashevich & M. B. Mostovski. 2002. Order Diptera Linné, 1758. The true flies (=Muscida Laicharting, 1781). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 227–240. Kluwer Academic Publishers: Dordrecht.
[B87] Boothroyd, I. K. G. 1987. Taxonomic composition and life cycles of Chironomidae (Diptera) in a northern New Zealand stream. Entomologica Scandinavica Supplement 29: 15–22.
[B04] Boothroyd, I. K. G. 2004. A new species of Naonella Boothroyd (Chironomidae: Orthocladiinae) from New Zealand. New Zealand Entomologist 27: 11–15.
[B72] Brundin, L. 1972. Phylogenetics and biogeography. Systematic Zoology 21 (1): 69–79.
[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).
[CS77] Cramp, S., & K. E. L. Simmons (eds) 1977. Handbook of the Birds of Europe, the Middle East and North Africa: The Birds of the Western Palaearctic vol. 1. Ostrich to Ducks. Oxford University Press: Oxford.
[F61] Freeman, P. 1961. The Chironomidae (Diptera) of Australia. Australian Journal of Zoology 9 (4): 611–735.
[G09] Gerecke, R. 2009. Revisional studies on the European species of the water mite genus Lebertia Neuman, 1880 (Acari: Hydrachnidia: Lebertiidae). Abhandlungen der Senckenberg Gesellschaft für Naturforschung 566: 1–144.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.
[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.
[N91] Norris, K. R. 1991. General biology. In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 68–108. Melbourne University Press: Carlton (Victoria).
[O81] Oliver, D. R. 1981. Chironomidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 423–458. Research Branch, Agriculture Canada.
[PH10] Pinder, A. M., S. A. Halse, R. J. Shiel & J. M. McRae. 2010. An arid zone awash with diversity: patterns in the distribution of aquatic invertebrates in the Pilbara region of Western Australia. Records of the Western Australian Museum Supplement 78 (1): 205–246.
[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.
[P94] Pugh, P. J. A. 1994. Non-indigenous Acari of Antarctica and the sub-Antarctic islands. Zoological Journal of the Linnean Society 110: 207–217.
[R86] Reiss, F. 1986. Erster Nachtrag zur Chironomidenfauna Bayerns (Diptera, Chironomidae). Spixiana 9 (2): 175–178.
[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.
[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
[S89] Skuse, F. A. A. 1889. Diptera of Australia. Part VI.—The Chironomidae. Proceedings of the Linnean Society of New South Wales, series 2, 4 (2): 215–311, pls 11–14.
[SA09] Spies, M., T. Andersen, J. H. Epler & C. N. Watson Jr. 2009. Chironomidae (non-biting midges). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 437–480. NRC Research Press: Ottawa.
[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).
[Z02] Zherikhin, V. V. 2002. Ecological history of the terrestrial insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 331–388. Kluwer Academic Publishers: Dordrecht.
There are nine subfamilies recognised, yet I see only six on your list. Where do you get your classification from?
The sources for each are indicated alongside the names. So far, I've just gotten most of them from more general references on insects; presumably, I haven't yet gone through references that would give me the other three. Hence, also, the taxa listed as just general Chironomidae at the end, which presumably would be placed more specifically once I've found and taken info from an appropriate source.