Belongs within: Eugnatha.
Contains: Neoatractosomatoidea, Anthroleucosomatidae, Cleidogonoidea, Diplomaragnidae, Conotylidae, Megalotylidae, Metopidiotrichinae.
The Chordeumatida are a group of relatively short, bristly millipedes. Gonopods are primitively derived from the modified eighth leg pair only, with the ninth pair modified into posterior gonopods in the Chordeumatidea and Heterochordeumatoidea (Shear 2003b).
Characters (Hoffman 1990): Body with 26–30 segments in adults; ocelli usually present; gnathochilarium with short basal mentum and elongate lingual lamellae; ozopores absent; metaterga with well-defined median longitudinal suture; sterna not firmly attached to adjoining sternites but more or less flexibly articulated by connective tissue; males with coxal sacs in ninth and tenth pairs of legs only; gonopods composed of both pairs of legs of seventh segment.
<==Chordeumatida [Chordeumida]
| i. s.: NeoatractosomatoideaM03
| AnthroleucosomatidaeM03
| Nanogona Cook 1895H00
|--ChordeumatideaS03b
| | i. s.: ChordeumatidaeM03
| | |--Chordeumella broelemanni (Verhoeff 1897)M03
| | `--Chordeuma silvestre [incl. Craspedosoma nemasomoides, Cr. polydesmoides]B82
| | Urochordeuma [Urochordeumatidae]H90
| `--CleidogonoideaH90
|--StriariideaS03b
| |--Caseyidae [Caseyoidea]S03b
| | |--CaseyaH90
| | |--OpionaH90
| | |--PlacernaH90
| | |--VasingtonaH90
| | `--ZantonaH90
| `--StriarioideaS03b
| |--Apterourus Loomis 1966 [Apterouridae]S03b
| | |--*A. horizontalis Loomis 1966S03b
| | `--A. serratus Shear 2003S03b
| `--+--RhiscosomidesH90 [RhiscosomididaeS03b]
| `--StriariidaeS03b
| |--VaferiaH90
| `--SpeostriariaH90
|--+--DiplomaragnidaeS99
| `--Heterochordeumatoidea [Heterochordeumatidea]S03b
| |--HeterochordeumatidaeS02a
| |--Adrityla [Adritylidae]H90
| |--ConotylidaeH90
| |--MegalotylidaeS02b
| `--Metopidiotrichidae [Metopidiothricidae]S02a
| |--Schedotrigona Silvestri 1903 [Schedotrigonidae, Schedotrigoninae]S02a
| |--MetopidiotrichinaeS02a
| `--Neocambrisomatinae [Neocambrisomatidae]S02a
| |--Neocambrisoma Mauriès 1987S02a
| |--Nesiothrix Shear & Mesibov 1997S02a
| |--Nipponothrix Shear & Tanabe 1994S02a
| `--Australeuma Golovatch 1987S02a
| `--A. mauriesi Shear & Mesibov 1997S02a
|--CraspedosomatideaMG00
| | i. s.: Craspedosoma [Craspedosomatidae, Craspedosomidae]MG00
| | |--C. affine Koch & Berendt 1854RB93
| | |--C. angulatum Koch & Berendt 1854RB93
| | |--C. armatum (Gerstfeldt 1859) (n. d.) [=Julus armatus]MG00
| | |--C. cylindricum Stuxberg 1876 (n. d.)MG00
| | `--C. simileA99
| `--MastigophorophylloideaM03
| |--MastigophorophyllidaeM03
| |--HoffmaneumatidaeM03
| `--AltajellidaeMG00
| |--Elongeuma Golovatch 1982MG00
| |--Altajella Gulička 1972MG00
| | `--*A. pallida Gulička 1972MG00
| `--Teleckophoron Gulička 1972MG00
| `--*T. montanum Gulička 1972MG00
`--Brannerioidea [Craspedosmatidea]SL03
|--NiponiosomaSH98 [NiponiosomatidaeSL03]
|--TrachygonidaeSL03
|--HeterolatzeliidaeSL03
|--ChaemosomatidaeSL03
|--Microlympia Shear & Leonard 2003 [Microlympiidae]SL03
| `--*M. echina Shear & Leonard 2003SL03
|--Branneria Bollman 1893 [Branneriidae]S03a
| |--*B. carinata (Bollman 1888) [=Craspedosoma carinata]S03a
| `--B. bonoculus Shear 2003S03a
|--TingupidaeSL03
| |--Tingupa Chamberlin 1910SL03
| | `--T. pallidaSH98
| |--Buotus Chamberlin 1940SL03
| | `--B. carolinusSH98
| `--Blancosoma Shear & Hubbard 1998SL03
| `--*B. scaturgo Shear & Hubbard 1998SH98
`--KashmireumatidaeS02b
|--KashmireumaS02b
| |--K. nepalensisS02b
| |--K. nielseniS02b
| `--K. schawallariS02b
`--Vieteuma Golovatch 1984S02b
|--*V. topaliS02b
|--V. hubeiensis Mauriés & Nguyen-Duy Jaquemin 1997S02b
`--V. longi Shear 2002S02b
*Type species of generic name indicated
References
[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (translated: 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).
[B82] Berlese, A. 1882. Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Fascicle 1. Typis Fratelli Salmin: Patavii.
[H00] Hannibal, J. T. 2000. Hexecontasoma, a new helminthomorph millipede (Hexecontasomatidae n. fam.) from the Mazon Creek, Illinois, fauna (Carboniferous, North America). Fragmenta Faunistica 43 (Suppl.): 19–35.
[H90] Hoffman, R. L. 1990. Diplopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 835–860. John Wiley & Sones: New York.
[M03] Mauriès, J.-P. 2003. Schizmohetera olympica sp. n. from Greece, with a reclassification of the superfamily Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta 12: 9–16.
[MG00] Mikhaljova, E. V., & S. I. Golovatch. 2000. A review of the millipede fauna of Siberia (Diplopoda). Arthropoda Selecta 9: 103–118.
[RB93] Ross, A. J., & D. E. G. Briggs. 1993. Arthropoda (Euthycarcinoidea and Myriapoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 357–361. Chapman & Hall: London.
[S99] Shear, W. A. 1999. The milliped genus Diplomaragna confirmed for Taiwan, with the description of a new species (Diplopoda, Chordeumatida, Diplomaragnidae). Myriapodologica 6 (2): 11–18.
[S02a] Shear, W. A. 2002a. The millipede genus Metopidiothrix Attems (Diplopoda: Chordeumatida: Metopidiotrichidae). Invertebrate Systematics 16: 849–892.
[S02b] Shear, W. A. 2002b. Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63–72.
[S03a] Shear, W. A. 2003a. Branneria bonoculus, n. sp., a second species in the North American milliped family Branneriidae (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa 233: 1–7.
[S03b] Shear, W. A. 2003b. A second species of the rare milliped family Apterouridae (Diplopoda: Chordeumatida: Striarioidea). Proceedings of the Biological Society of Washington 116 (2): 509–514.
[SH98] Shear, W. A., & D. A. Hubbard, Jr. 1998. Cave millipeds of the United States. IV. A new genus and species from a high altitude cave in Colorado (Diplopoda, Chordeumatida, Tingupidae). Myriapodologica 5 (8): 85–94.
[SL03] Shear, W. A., & W. P. Leonard. 2003. Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa 243: 1–11.