Clusiidae

Belongs within: Schizophora.

The Clusiidae are a group of relatively slender flies that are commonly associated with humid habitats. They are characterised by the possession of a porrect antenna with a dorsoapical arista and triangular extensions on the outer and sometimes inner distal margins of the pedicel. In most members of the subfamily Clusiodinae, the anterior pair of fronto-orbital bristles is reclinate whereas members of other subfamilies, and of the clusiodine genus Heteromeringia, have these bristles inclinate (Lonsdale & Marshall 2010).

Characters (from Lonsdale & Marshall 2010): Relatively elongate flies. Colour various but usually all or partly yellow with wing frequently infuscated. Arista pubescent to densely plumose; vibrissa well developed; face usually soft, receding. Outer and sometimes inner distal margins of pedicel with triangular extensions, outer extension distinct and acute in Clusiinae and Sobarocephalinae, blunt and obtuse in Clusiodinae. Ocellar and/or postocellar bristles usually small to absent, divergent when present. Subcosta complete and subcostal break present (sometimes indistinct in Clusiodinae).

<==Clusiidae [Clusiodidae, Heteroneuridae]
    |--ClusiinaeLM10
    |    |--Clusia Haliday 1838G05
    |    |    |--C. flava (Meigen 1830)G05
    |    |    `--C. lateralisS87
    |    |--PhylloclusiaLM10
    |    |--Tetrameringia ustulataLM10, O98
    |    |--MelanoclusiaLM10
    |    `--ApiochaetaLM10
    |         |--A. bicolorLM10
    |         `--A. connexaLM10
    |--SobarocephalinaeLM10
    |    |--Chaetoclusia bakeriLM10
    |    |--Procerosoma aliniLM10
    |    `--SobarocephalaLM10
    |         |--S. beckeri Hennig 1956S63b
    |         |--S. boliviana Soós 1963S63b
    |         |--S. columbiensis Soós 1962S63b
    |         |--S. dudichi Soós 1963S63b
    |         |--S. festiva Czerny 1929S63b
    |         |--S. flavaLM10
    |         |--S. flavisetaLM10
    |         |--S. fumipennis Soós 1963S63b
    |         |--S. lanei Curran 1939S63b
    |         |--S. latifronsLM10
    |         |--S. nigrofacies Soós 1962S63b
    |         |--S. nitida Soós 1963S63b
    |         |--S. picta Hennig 1938S63b
    |         |--S. pictipennis (Kertész 1903) [=Monorrhexa pictipennis]S63b
    |         |--S. plumata Melander & Argo 1925S63b
    |         |--S. pruinosa Soós 1962S63b
    |         |--S. quadrimaculata Soós 1963S63b
    |         |--S. sabroskyi Soós 1963S63b
    |         |--S. strigata Hennig 1938S63b
    |         |--S. styskali Soós 1963S63b
    |         |--S. testacea [incl. Chaetoclusia affinis]S87
    |         |--S. variegata Melander & Argo 1925S63b
    |         `--S. zeugma Hennig 1938S63b
    `--ClusiodinaeLM10
         |--Craspedochaeta concinnaLM10
         |--Clusiodes Coquilett 1904G05
         |    |  i. s.: C. johnsoniLM10
         |    |--C. (Clusiodes)G05
         |    |    |--C. (C.) albimanus (Meigen 1830)G05
         |    |    |--C. (C.) caledonicus (Collin 1912)G05
         |    |    `--C. (C.) gentilis (Collin 1912)G05
         |    |--C. (Clusiaria)G05
         |    |    |--C. (C.) apicalis (Zetterstedt 1848)G05
         |    |    |--C. (C.) freyi Tuomikoski 1933G05
         |    |    |--C. (C.) geomyzinus (Fallén 1823) [=Heteroneura geomyzina]G05
         |    |    |--C. (C.) pictipes (Zetterstedt 1855)G05
         |    |    `--C. (C.) ruficollis (Meigen 1830) [incl. C. fascialis (Collin 1953)]G05
         |    `--C. (Columbiella) verticalis (Collin 1912)G05
         |--Hendelia Czerny 1903LM10, G05
         |    |--*H. beckeri Czerny 1903LM10, S63a [incl. H. nigripalpis Czerny 1928S63a]
         |    |--H. kinetrolicrosLM10
         |    `--H. mirabilis [=Clusiodes mirabilis]LM10
         |--Heteromeringia [incl. Tranomeringia]LM10
         |    |--H. czernyiLM10
         |    |--H. fumipennisLM10
         |    |--H. nigrimanaG05
         |    |--H. nitidaLM10
         |    `--H. norrisiCM91
         `--AllometoponLM10
Clusiidae incertae sedis:
  ElectroclusiodesLM10
    |--E. meunieriS87
    `--E. radiospinosaS87
  TrichoclusiaM87
  ChaetoclusiellaM87
  Prohendelia Frey 1960S63a
    |--P. (Prohendelia)S63a
    |    |--*P. (P.) orientalis Frey 1960S63a
    |    `--P. (P.) freyi Soós 1963S63a
    `--P. (Aristohendelia) orientalisS63a
  Parahendelia McAlpine 1960S63a
    `--*P. latifrons McAlpine 1960S63a
  Paraclusia Czerny 1902G05
    `--P. tigrina (Fallén 1820) [=Heteromyza tigrina]G05

*Type species of generic name indicated

References

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[G05] Greve, L. 2005. Atlas of the Clusiidae (Diptera) in Norway. Insecta Norvegiae 7: 1–27.

[LM10] Lonsdale, O., & S. A. Marshall. 2010. Clusiidae (clusiid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1041–1048. NRC Research Press: Ottawa.

[M87] McAlpine, J. F. 1987. Acartophthalmidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 859–861. Research Branch, Agriculture Canada.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[S63a] Soós, Á. 1963a. Prohendelia (Prohendelia) freyi spec. nov., and some notes on the genus Hendelia Czerny s. l. (Diptera: Clusiidae). Acta Zoologica Academiae Scientiarum Hungaricae 9 (1–2): 177–182.

[S63b] Soós, Á. 1963b. Identification key to the species of the “plumata-group” of the genus Sobarocephala Czerny (Diptera: Clusiidae). Acta Zoologica Academiae Scientiarum Hungaricae 9 (3–4): 391–396.

[S87] Soós, Á. 1987. Clusiidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 853–857. Research Branch, Agriculture Canada.