Belongs within: Chrysomelidae.
Contains: Clytrasoma.
The Clytrini are a group of leaf beetles whose larvae are commonly found in association with ant nests. Members of this group have a kotpresse usually composed of five sclerites with a flat central plate accompanied by two lateral sclerites with broadened horizontal terminations, and two well separated ventral sclerites with large apodemes (Schöller & Witte 2007).
<==ClytriniSW07 |--Ischiopachys Chevrolat 1836 [Ischiopachina]RC02 | `--I. proteusRC02 |--MegalostominaRC02 | |--Euryscopa Lacordaire 1848RC02 | |--Coleorozena Moldenke 1981RC02 | |--Coscinoptera Lacordaire 1848RC02 | | `--C. aeneipennis (LeConte 1858)RC02 | |--Coleothorpa Moldenke 1981RC02 | | `--C. dominicana (Fabricius 1801) [=Coscinoptera dominicana]SW07 | `--Megalostomis Chevrolat 1836RC02 | |--M. (Coleobyersa Moldenke 1981)RC02 | |--M. (Heterostomis Lacordaire 1848)RC02 | |--M. (Minturnia Lacordaire 1848)RC02 | |--M. (Pygidiocarina Moldenke 1970)RC02 | |--M. (Scaphigenia Lacordaire 1848)RC02 | `--M. (Snellingia Moldenke 1981)RC02 |--BabiinaRC02 | |--Saxinis Lacordaire 1848RC02 | | `--S. (Boreosaxinis Moldenke 1981)RC02 | | |--S. (B.) omogera Lacordaire 1848RC02 | | `--S. (B.) saucia LeConte 1857RC02 | |--Babia Chevrolat 1836 [incl. Harpasta Gistel 1847]RC02 | | |--B. (Babia) quadriguttata Olivier 1796RC02 | | |--B. (Archaebabia Moldenke 1981)RC02 | | | |--B. (A.) costalisdebaja Moldenke 1970RC02 | | | `--B. (A.) tetraspilota LeConte 1858RC02 | | |--B. (Coleolacordairei Moldenke 1981)RC02 | | |--B. (Heterobabia Monrós 1951)RC02 | | `--B. (Megababia Moldenke 1981)RC02 | `--Urodera Lacordaire 1848RC02 | |--U. (Austrurodera Moldenke 1981)RC02 | |--U. (Boreurodera Moldenke 1981)RC02 | | |--U. (B.) cruciferaRC02 | | | |--U. c. cruciferaRC02 | | | `--U. c. texana Schaeffer 1919RC02 | | `--U. (B.) dilaticollis Jacoby 1889RC02 | |--U. (Familiurodera Moldenke 1981)RC02 | `--U. (Stereomoides Moldenke 1981)RC02 `--ClytrinaRC02 | i. s.: Anomoea Agassiz 1846 [=Anomoia Chevrolat 1836 non Walker 1835; incl. Euthyna Gistel 1847]RC02 | `--A. laticlavia (Forster 1771)RC02 |--ClytraSW07 [incl. AntipusL02; Clytrina] | | i. s.: C. annamita Lefèvre 1889L89a | | C. atraphaxidisR26 | | C. bimaculataR26 | | C. concolorR26 | | C. duodecimmaculata (see below for synonymy)L89a | | C. laeviuscula (Ratzeburg 1837)SW07 | | C. longipesL02 | `--C. (Clytra)B14 | |--C. (C.) alienaB14 | `--C. (C.) quadripunctata (Linnaeus 1758)B14 [=Cryptocephalus (Clythra) quadripunctatusG20] `--+--+--Tituboea sexmaculata (Fabricius 1781)SW07 | `--+--Lachnaia paradoxa (Olivier 1808)SW07 | `--+--Macrolenes Chevrolat 1837SW07 | | `--M. dentipes (Olivier 1808)SW07 | `--Labidostomis Germar 1817SW07 | |--L. bipunctata (Mannerheim 1825)SW07 | |--L. heteromalla Lefèvre 1889L89c | |--L. pallidipennisL89c | `--L. sibiricaC01 `--+--ClytrasomaSW07 `--Smaragdina Chevrolat 1836SW07, RC02 | i. s.: S. limbata (Stéven 1806)SW07 |--S. (Smaragdina) (see below for synonymy)RC02 | |--‘Gynandrophthalma’ cochinchinensis Lefèvre 1889L89a | |--‘Gynandrophthalma’ divisa Jacoby 1889J89 | |--‘Gynandrophthalma’ malayana Baly 1865L89a | |--S. (S.) militaris LeConte 1858RC02 | | |--S. m. militarisRC02 | | `--S. m. arizonica Schaeffer 1919RC02 | `--‘Gynandrophthalma’ sexmaculata Jacoby 1889J89 `--S. (Otiocephala Lefèvre 1872)RC02 Clytrini incertae sedis: Diapericera Lacordaire 1848 [Eoclytrina]SW07 DiapromorphaL89a |--D. bifasciata (Illig. 1800) [=Clytra bifasciata; incl. C. bicolor Weber 1801, Aspidolopha buqueti]L89a |--D. melanophthalma [=Aspidolopha melanophthalma]J89 |--D. melanopusJ89 |--D. pinguis [incl. Titubaea laportei Baly 1865]L89a `--D. sodalis [=Aetheomorpha sodalis]L89a DachrysL90 |--D. monogrammaL90 `--D. pumilioL90 CoptocephalaSW07 |--C. jacobyi Lefèvre 1889L89b `--C. unifasciata (Scopoli 1763)SW07 [incl. C. unifasciata var. gebleriC01]
Clytra duodecimmaculata [=Cryptocephalus duodecimmaculatus; incl. Chrysomela cryptocephala Hornstedt 1788, Cr. javanus]L89a
Smaragdina Chevrolat 1836 (Smaragdina) [incl. Calyptorhina Lacordaire 1848, Cyaniris Chevrolat 1836 non Dalman 1816, Carmentis Gistel 1847, Exomis Weise 1889, Gynandrophthalma Lacordaire 1848, Monrosia Medvedev 1971, Necyomantes Gistel 1847, Smaragdinella Medvedev 1971 non Adams 1848]RC02
*Type species of generic name indicated
References
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[J89] Jacoby, M. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVII.—List of the phytophagous Coleoptera obtained by Signor L. Fea at Burmah and Tenasserim, with descriptions of the new species. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 147–237.
[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[L89a] Lefèvre, É. 1889a. Contribution a la faune Indo-Chinoise. 4e mémoire. Cryptocéphalides, clytrides et eumolpides. Annales de la Société Entomologique de France, 6e série 9: 287–299.
[L89b] Lefèvre, É. 1889b. Description suivante d’une nouvelle espèce de clytrides, du genre Coptocephala, provenant de l’Afrique tropicale orientale. Annales de la Société Entomologique de France, 6e série 9: lxx–lxxi.
[L89c] Lefèvre, É. 1889c. Description d’une nouvelle espèce de clytrides. Annales de la Société Entomologique de France, 6e série 9: cxix.
[L90] Lefèvre, É. 1890. Voyage de M. E. Simon au Venezuela (Décembre 1887–Avril 1888). 6e mémoire. Clytrides, lamprosomides et eumolpides. Annales de la Société Entomologique de France, 6e série 9: 329–336.
[RC02] Riley, E. G., S. M. Clark, R. W. Flowers & A. J. Gilbert. 2002. Chrysomelidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 617–691. CRC Press.
[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
[SW07] Schöller, M., & V. Witte. 2007. A review of the genus Clytrasoma Jacoby 1908, with description of a new species collected within a Camponotus sp. ant-nest (Insecta: Coleoptera, Chrysomelidae; Hymenoptera, Formicidae). Senckenbergiana Biologica 87 (1): 51–61.