Coenosiinae

Belongs within: Muscidae.
Contains: Lispini, Limnophora, Coenosiini.

The Coenosiinae are a group of muscid flies with carnivorous larvae adapted for aquatic habitats. Members include the cosmopolitan genus Spilogona, characterised by the basisternum of the prosternum and the posterior surface of the hind coxa being bare, and the antenna being shortly pubescent with the hairs no longer than the basal diameter of the arista. Paralimnophora is a genus endemic to New Zealand in which the face has a high gena, about half the height of the eye (Couri 2010). Exsul, another New Zealand endemic genus, has remarkable broad wings, about as wide as long (Couri 2010).

<==Coenosiinae [Limnophorini, Lispinae]KP10
    |--+--LispiniKP10
    |  `--LimnophoraKP10
    `--+--CoenosiiniKP10
       `--+--Villeneuvia aestuumKP10
          `--Spilogona Schnabl 1911KP10, F92 [incl. MydaeinaHV87]
               |--*S. carbonella (Zetterstedt 1845) [=Aricia carbonella, Limnophora (*Spilogona) carbonella]F92
               |--S. almqvistii (Holmgren 1880) [=Aricia almqvistii]F92
               |--S. anthrax (Bigot 1885) [=Limnophora anthrax]H72
               |--S. caliginosaKP10
               |--S. costalis (Stein 1907) [=Limnophora costalis]F92
               |--S. disparKP10
               |--S. leptocerci Mou in Fan 1992F92
               |--S. litorea (Fallen 1823) [=Musca litorea]F92
               |    |--S. l. litoreaF92
               |    `--S. l. yaluensis Ma & Wang in Fan 1992F92
               |--S. littoralis Mou in Fan 1992F92
               |--S. obscuraHV87
               |--S. setigera (Stein 1907) [=Limnophora setigera]F92
               |--S. spinicosta (Stein 1907) [=Limnophora spinicosta]F92
               |--S. surda (Zetterstedt 1845) [=Aricia surda]F92
               `--S. taheensis Ma & Cui in Fan 1992F92
Coenosiinae incertae sedis:
  Heliographa Malloch 1921F92
    `--‘Hydrotaea’ javana Macquart 1851 [incl. Limnophora tonsa Stein 1909, *Heliographa intonsa]C10
  Pseudolimnophora Strobl 1893F92
  Neolimnophora Schnabl & Dziedzicki 1911F92
  Exsul Hutton 1901C10
    `--*E. singularis Hutton 1901C10
  Limnohelina Malloch 1930C10
    `--*L. debilis [=Cordylura debilis]C10
  Paracoenosia Malloch 1938C10
    `--*P. tonnoiri Malloch 1938C10
  Paralimnophora Lamb 1909C10
    `--*P. depressa Lamb 1909C10

*Type species of generic name indicated

References

[C10] Couri, M. S. 2010. Key to the Australasian and Oceanian genera of Muscidae (Diptera). Revista Brasileira de Entomologia 54 (4): 529–544.

[F92] Fan Z. 1992. Key to the Common Flies of China 2^nd ed. Science Press: Beijing.

[H72] Huckett, H. C. 1972. Notes on Bigot’s North American type-specimens at the University Museum, Oxford (Diptera: Anthomyiidae, Muscidae). Pan-Pacific Entomologist 48 (2): 81–85.

[HV87] Huckett, H. C., & J. R. Vockeroth. 1987. Muscidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1115–1131. Research Branch, Agriculture Canada.

[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine’s fly. Systematic Entomology 35: 614–635.