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Belongs within: Cucujoidea.
The Cryptophilinae are a group of saprophagous or mycophagous beetles mostly found amongst leaf litter (Leschen 2003).
Characters (from Leschen 2003): Mandible usually with subapical serrations; lacinia with one or no lacinial spines; mentum lacking lateral pockets, usually with medial carina; apical labial palpomere as wide as or wider than long, usually not securiform; ligular membrane as wide or wider than prementum; transverse gular line usually present as shallow groove (rarely absent or deep); antennal insertion hidden in dorsal view; supraocular line usually present to level above eye; vertex of head commonly with stridulatory file; pronotum usually not strongly constricted at base; anterior angles of pronotum poorly or well developed; anterior margin of pronotum usually serrate; pronotal pits usually present; procoxal cavity internally closed, usually open behind or closed by hypomeron, edge of hypomeron usually smooth; mesometaventral articulation, if present, dicondylic; width of mesoventral process usually less than coxa; submesocoxal lines usually absent; abdominal ventrites 1 and 2 usually free; abdominal calli present; metafurcal lamina usually absent with median stalk of metendosternite poorly or well developed; spermatheca rounded or elongate; apical pit of spermatheca usually absent; accessory gland of spermatheca usually present; tarsomere 1 usually longer than tarsomere 2 and 3; tarsomere 4 usually exposed in ventral view; tarsal shelf of tarsomere 5 usually absent; empodium with one or two setae; elytral punctation striate or confused with scutellary striole absent; elytra usually narrowly explanate; hind with with radial cell and wedge cell absent.
<==Cryptophilinae [Cryptophilidae] |--EmpocryptiniL03 | |--Lobosternum clavicornaL03 | `--+--Lepidotoramus grouvelleiL03 | `--Empocryptus Sharp 1900L03 | `--E. ovalisL03 `--+--Toramini [Toraminae]L03 | |--+--Loberoschema bimaculataL03 | | `--StengitaL03 | `--+--AtomaropsL03 | | |--A. curvitibialisL03 | | `--A. lewisiL03 | `--Toramus Grouvelle 1916L03, LS02 [incl. Itomarus Reitter 1919LS02, Tomarus LeConte 1861 non Erichson 1847LS02] | |--‘Tomarops’ bimaculatusL03 | |--T. hirtellusL03 | |--‘Tomarops’ longitarsisL03 | `--T. pulchellusRS15 `--Cryptophilini [Xenoscelinini]L03 | i. s.: Chinophagus Ljubarsky 1997L03 |--Brachypterosa peckorumL03 `--+--+--Crowsonguptus mexicanusL03 | `--Cryptophilus Reitter 1874 [incl. Tomarops Grouvelle 1903]L03 | |--*C. integer (Heer 1841) [=Cryptophagus integer]L03 | |--C. allotrius [=Cryptophagops allotrius]L03 | |--C. fulminalisLS02 | |--C. leonensis [=Cryptophagops leonensis]L03 | |--C. mnionomoides [=Cryptophagops mniomonoides]L03 | `--‘*Tomarops’ punctatus Grouvelle 1903L03 `--+--Cathartocryptus Sharp 1886 [incl. Xenoscelinus Grouvelle 1910]L03 | |--C. maculosus (Broun 1881) (see below for synonymy)L03 | `--‘*Xenoscelinus’ malaicus Grouvelle 1910L03 `--LoberopsyllusL03 |--L. explanatus Leschen & Ashe 1999B14 |--L. halffteriB14 |--L. oculatusL03 `--L. traubiB14
Cathartocryptus maculosus (Broun 1881) [=Paramecosoma maculosa, Xenoscelinus maculosus; incl. *C. obscurus Sharp 1886]L03
*Type species of generic name indicated
References
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).
[L03] Leschen, R. A. B. 2003. Erotylidae (Insecta: Coleoptera: Cucujoidea): phylogeny and review. Fauna of New Zealand 47: 1–108.
[LS02] Leschen, R. A. B., & P. E. Skelley. 2002. Languriidae Wiedeman1823. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 343–347. CRC Press.
[RS15] Robertson, J. A., A. Ślipiński, M. Moulton, F. W. Shockley, A. Giorgi, N. P. Lord, D. D. McKenna, W. Tomaszewska, J. Forrester, K. B. Miller, M. F. Whiting & J. V. McHugh. 2015. Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia). Systematic Entomology 40: 745–778.