Cyrtospiriferoidea

Partial brachial valve of Cyrtiopsis animasensis, from here.

Belongs within: Spiriferida.
Contains: Cyrtospirifer.

The Cyrtospiriferoidea are a group of Devonian spiriferids characterised by the presence of a delthyrial plate. The shell is biconvex and generally transverse with a median fold and sulcus.

<==CyrtospiriferoideaRG04
|--Spinocyrtiidae [Guerichellinae, Spinocyrtiinae]RG04
| |--Acutatheca Stainbrook 1945BJ65
| | `--*A. propria Stainbrook 1945BJ65
| |--Alatiformia Struve 1963M65
| | `--*A. alatiformis (Drevermann 1907) [=Spirifer alatiformis]M65
| |--Allanella Crickmay 1953 [=Allanaria Crickmay 1953]BJ65
| | `--*A. allani (Warren 1944) [=Spirifer allani, *Allanaria allani]BJ65
| |--Chimaerothyris Paulus, Struve & Wolfart 1963M65
| | `--*C. hotzi Paulus, Struve & Wolfart 1963M65
| |--Eosyringothyris Stainbrook 1943BJ65
| | |--*E. aspera (Hall 1858) [=Spirifera aspera]BJ65
| | `--E. calviniMB12
| |--Subcuspidella Mittmeyer 1965 [incl. Tenuicostella]RG04
| | |--*S. subcuspidata (Schnur 1851) [=Spirifer subcuspidatus]RG04
| | |--‘Tenuicostella’ comellanensisMB12
| | `--S. incerta (Fuchs 1909)RG04
| |--Adolfia Gürich 1909 [=Guerichella Paeckelmann 1931, Guerichia (l. c.); incl. Plectospirifer Grabau 1931]BJ65
| | |--*A. deflexa (Roemer 1843) [=Spirifer deflexus, *Guerichella deflexa]BJ65
| | |--‘Guerichella’ angustistellataBJ65
| | |--A. bifidaBJ65
| | |--‘Spirifer (*Plectospirifer)’ heimi Grabau 1931BJ65
| | `--A. talasicaH79
| `--Spinocyrtia Frederiks 1916 [incl. Platyrachella Fenton & Fenton 1924]BJ65
| |--*S. granulosa (Conrad 1839) [=Delthyris granulosa]BJ65
| |--S. affinis (Fuchs 1929)HB93
| |--S. clintoniMB12
| |--S. iowensisMB12
| |--S. macbridei (Calvin 1883) [=Spirifera macbridei, *Platyrachella macbridei]BJ65
| |--‘Platyrachella’ mesastralisN79
| |--S. mouranteMB12
| `--S. struniana (Gosselet 1879)HB93
`--Cyrtospiriferidae [Cyrtospiriferinae]RG04
|--CyrtospiriferRG04
|--Tipispirifer oppilatus Grant 1976HB93
|--Platyspirifer Grabau 1931BJ65
| `--*P. paronai (Martelli 1902) [=Schizophoria paronai, Theodossia (*Platyspirifer) paronai]BJ65
|--Tenticospirifer Tien 1938BJ65
| `--*T. tenticulum (de Verneuil 1845) [=Spirifer tenticulum]BJ65
|--Arctospirifer Stainbrook 1950BJ65
| `--*A. constrictus Stainbrook 1950BJ65
|--Austrospirifer Glenister 1955BJ65
| `--*A. variabilis Glenister 1955BJ65
|--Dmitria Sidyachenko 1961BJ65
| `--*D. romanowskii (Nalivkin 1930) [=Spirifer (Cyrtospirifer) romanowskii]BJ65
|--Prospira Maxwell 1954BJ65
| `--*P. typa Maxwell 1954BJ65
|--Schizospirifer Grabau 1931BJ65
| `--*S. latistriatus (Frech 1911) [=Spirifer aperturatus var. latistriatus]BJ65
|--Sphenospira Cooper 1954BJ65
| `--*S. alta (Hall 1866) [=Spirifera alta]BJ65
|--Sulcatospirifer Maxwell 1954BJ65
| `--*S. primus Maxwell 1954BJ65
|--Syringospira Kindle 1909BJ65
| `--*S. prima Kindle 1909BJ65
|--Eodmitria supradisjunctus (Obrutchew 1917)HB93
| |--E. s. supradisjunctusHB93
| `--E. s. boloniensis Brice 1982HB93
|--Indospirifer Grabau 1931BJ65
| |--*I. padaukpinensis (Reed 1908) [=Spirifer padaukpinensis]BJ65
| | |--I. p. padaukpinensisBJ65
| | `--I. p. maoerhchuanensisBJ65
| `--I. pseudowilliamsiHB93
`--Cyrtiopsis Grabau 1923 (see below for synonymy)BJ65
|--*C. davidsoni Grabau 1923BJ65
|--C. animasensisN79
|--C. graciosa Grabau 1923 [=C. (*Grabauicyrtiopsis) graciosa]BJ65
|--C. intermediaBJ65
|--‘Uchtospirifer’ murchisonianusH79
|--*Uchtospirifer’ nalivkini Lyashenko 1957BJ65
`--C. transversa Grabau 1931 [=C. (*Sinocyrtiopsis) transversa]BJ65

Cyrtiopsis Grabau 1923 [incl. Alphacyrtiopsis Gatinaud 1949 (nom. vet.), Betacyrtiopsis Gatinaud 1949 (nom. vet.), Grabauicyrtiopsis Gatinaud 1949, Paracyrtiopsis Gatinaud 1949 (nom. vet.), Sinocyrtiopsis Gatinaud 1949, Uchtospirifer Lyashenko 1957]BJ65

*Type species of generic name indicated

References

[BJ65] Boucot, A. J., J. G. Johnson, C. W. Pitrat & R. D. Staton. 1965. Spiriferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 2 pp. H632–H728. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

[HB93] Harper, D. A. T., C. H. C. Brunton, L. R. M. Cocks, P. Copper, E. N. Doyle, A. L. Jeffrey, E. F. Owen, M. A. Parkes, L. E. Popov & C. D. Prosser. 1993. Brachiopoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 427–462. Chapman & Hall: London.

[H79] House, M. R. 1979. Devonian in the Eastern Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A183–A217. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[MB12] Mistiaen, B., D. Brice, M. K. Zapalski & C. Loones. 2012. Brachiopods and their auloporid epibionts in the Devonian of Boulonnais (France): comparison with other associations globally. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 159–188. Springer.

[M65] Moore, R. C. (ed.) 1965. Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 2. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).

[N79] Norris, A. W. 1979. Devonian in the Western Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A218–A253. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[RG04] Racheboeuf, P. R., R. Gourvennec, M. Deynoux & D. Brice. 2004. The Devonian of the Hodh area (Islamic Republic of Mauritania): paleontology and stratigraphy. Journal of Paleontology 78 (1): 98–110.

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