Belongs within: Neobatrachia.
Contains: Microhylidae, Laurentobatrachia, Saukrobatrachia, Pyxicephalidae, Ptychadenidae.
The Diplasiocoela are a clade of frogs supported by molecular analysis with many basal lineages having an African distribution (Feng et al. 2017). The basal Sooglossidae are small to medium-sized frogs with modified arciferal pectoral girdles that are endemic to the Seychelles (Nussbaum & Wu 2007).
<==DiplasiocoelaFB17
|--+--Nasikabatrachus Biju & Bossuyt 2003FG06 [NasikabatrachidaePW11]
| | `--*N. sahyadrensis Biju & Bossuyt 2003BB03
| `--Sooglossidae [Sooglossinae]PW11
| |--Sooglossus Boulenger 1906FB17, NW07 [incl. Nesomantis Boulenger 1909NW07]
| | |--*S. sechellensis (Boettger 1896) [=Arthroleptis sechellensis]NW07
| | `--S. thomasseti (Boulenger 1909) [=*Nesomantis thomasseti]NW07
| `--Sechellophryne Nussbaum & Wu 2007NW07
| |--*S. gardineri (Boulenger 1911) [=Nectophryne gardineri, Sooglossus gardineri]NW07
| `--S. pipilodryas (Gerlach & Willi 2002) [=Sooglossus pipilodryas]NW07
`--Ranoidea [Allodapanura, Firmisternia, Microhyloidae, Ranoides]FB17
| i. s.: Thamastosaurus gezeiFB17
|--MicrohylidaeFB17
`--+--Afrobatrachia [Brevicipitoidae]FB17
| |--LaurentobatrachiaPW11
| `--XenosyneunitanuraFG06
| |--Hemisus Günther 1859FG06 [Hemisidae, HemisotidaePW11, Hemisotina]
| | `--H. marmoratus Peters 1854DE08
| `--Brevicipitidae [Brevicipitina, Brevicipitinae]PW11
| | i. s.: Balebreviceps Largen & Drewes 1989FG06
| | Engystoma Fitzinger 1826 [Engystomidae]FG06
| |--Breviceps Merrem 1820FG06
| | | i. s.: B. macropsFB17
| | |--B. fuscusPW11
| | `--+--B. fichusPW11
| | `--B. mossambicusPW11
| `--+--Spelaeophryne Ahl 1924FB17, FG06
| | `--S. methneriPW11
| `--+--Callulina Nieden 1911FB17, FG06
| | |--C. kisiwamsituPW11
| | |--C. krefftiPW11
| | `--C. laphamiFB17
| `--Probreviceps Parker 1931FB17, FG06
| |--P. durirostrisPW11
| `--+--P. macrodactylusPW11
| `--P. uluguruensisPW11
`--Natatanura [Africanura, Ametrobatrachia, Ranoidae, Telmatobatrachia, Victoranura]FB17
|--SaukrobatrachiaFB17
`--+--PyxicephaloideaFG06
| |--PyxicephalidaeFB17
| `--+--Conraua Nieden 1908FG06 [ConrauidaeFB17, Conrauinae, Conrauini]
| | |--C. robustaPW11
| | `--+--C. crassipesPW11
| | `--C. goliathPW11
| `--Petropedetidae [Petropedetinae]FB17
| |--Arthroleptides Nieden 1911FG06
| | `--A. yakusiniFG06
| `--Petropedetes Reichenow 1874FG06
| | i. s.: P. euskircheniFB17
| | P. johnstoniFG06
| | P. natatorFG06
| |--+--P. martiensseniPW11
| | `--P. yakusiniPW11
| `--+--P. palmipesPW11
| `--+--P. cameronensisPW11
| `--+--P. newtoniPW11
| `--P. parkeriPW11
`--+--Odontobatrachus [Odontobatrachidae]FB17
| `--O. natatorFB17
`--+--PtychadenidaeFB17
`--Phrynobatrachidae [Phrynobatrachinae]FB17
|--Hemimantis Peters 1863 [Hemimantidae]FG06
`--Phrynobatrachus Günther 1862 (see below for synonymy)FG06
| i. s.: P. guineensisPM09
|--+--P. sandersoni (Parker 1935) [=Phrynodon sandersoni]FG06
| `--+--P. dendrobatesPW11
| `--P. krefftiiPW11
`--+--+--P. mababiensisPW11
| `--+--P. calcaratusPW11
| `--+--P. disparPW11
| `--P. levelevePW11
`--+--+--P. auritusPW11
| `--P. natalensisPW11
`--+--P. acridoidesPW11
`--+--P. africanus (Hallowell 1858)PW11, FG06 [=Dimorphognathus africanusFG06]
`--P. cricogasterPW11
Phrynobatrachus Günther 1862 [incl. Dimorphognathus Boulenger 1906, Phrynodon Parker 1935]FG06
*Type species of generic name indicated
References
[BB03] Biju, S. D., & F. Bossuyt. 2003. New frog family from India reveals an ancient biogeographical link with the Seychelles. Nature 425: 711–714.
[DE08] Doughty, P., & D. Edwards. 2008. A new species of Arenophryne (Anura: Myobatrachidae) from the central coast of Western Australia. Records of the Western Australian Museum 24 (2): 121–131.
[FB17] Feng, Y.-J., D. C. Blackburn, D. Liang, D. M. Hillis, D. B. Wake, D. C. Cannatella & P. Zhang. 2017. Phylogenomics reveals rapid, simultaneous diversification of three major clades of Gondwanan frogs at the Cretaceous–Paleogene boundary. Proceedings of the National Academy of Sciences of the USA 114 (29): E5864–E5870.
[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.
[NW07] Nussbaum, R. A., & S.-H. Wu. 2007. Morphological assessments and phylogenetic relationships of the Seychellean frogs of the family Sooglossidae (Amphibia: Anura). Zoological Studies 46 (3): 322–335.
[PW11] Pyron, R. A., & J. J. Wiens. 2011. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61: 543–583.