Ellesmeroceratidae

Belongs within: Nautilosiphonata.

The Lower Ordovician Ellesmeroceratidae were a family of straight-shelled or cyrtoconic cephalopods, many of them relatively small.

Characters (from Furnish & Glenister 1964): Orthoconic longicones and compressed endogastric breviconic cyrtocones with marginal siphuncle, ellipochoanitic septal necks and thick, layered connecting rings; often with irregular diaphragms.

<==Ellesmeroceratidae [Ellesmereoceratidae, Stemtonoceratidae, Robsonoceratidae]
    |--Paradakeoceras minor Flower 1964KE19, EK12
    |--Botellusoceras torpenseM13, KE19
    |--Eburnoceras pissinum Chen & Qi 1980K93
    |--Bridgeoceras Ulrich, Foerste & Miller 1943FG64
    |    `--*B. subannulatum Ulrich, Foerste & Miller 1943FG64
    |--Chepuloceras Unklesbay & Young 1956FG64
    |    `--*C. inelegans Unklesbay & Young 1956FG64
    |--Copiceras Ulrich, Foerste et al. 1944 [=Cophiceras (l. c.)]FG64
    |    `--C. erectum Ulrich, Foerste et al. 1944FG64
    |--Desioceras Cecioni 1953FG64
    |    `--*D. floweri Cecioni 1953FG64
    |--Hemichoanella Teichert & Glenister 1954FG64
    |    `--*H. canningi Teichert & Glenister 1954FG64
    |--Lebetoceras Teichert & Glenister 1954FG64
    |    `--*L. oepiki Teichert & Glenister 1954FG64
    |--Loxochoanella Teichert & Glenister 1954FG64
    |    `--*L. warburtoni Teichert & Glenister 1954FG64
    |--Oelandoceras Foerste 1932FG64
    |    `--*O. haelluddenense Foerste 1932FG64
    |--Paracyclostomiceras Cecioni 1953FG64
    |    `--*P. floweri Cecioni 1953FG64
    |--Quebecoceras Foerste 1925FG64
    |    `--*Q. quebecense (Whiteaves 1898) [=Cyrtoceras quebecense]FG64
    |--Smithvilloceras Ulrich, Foerste & Miller 1943FG64
    |    `--*S. thompsoni Ulrich, Foerste & Miller 1943FG64
    |--Stemtonoceras Ulrich & Foerste 1936FG64
    |    `--*S. elongatum Ulrich & Foerste 1936FG64
    |--Ventroloboceras Teichert & Glenister 1954FG64
    |    `--*V. furcillatum Teichert & Glenister 1954FG64
    |--Dongshanoceras Zou & Chen 1979CT83
    |    `--D. magnitubulatum Zou & Chen 1979CT83
    |--Eoectenolites Chen & Qi 1979CT83
    |    `--*E. suxianensis Chen & Qi in Chen et al. 1979CT83
    |--Hunyuanoceras Chen & Teichert 1983CT83
    |    `--*H. shanxiense Chen & Teichert 1983CT83
    |--Pseudendoceras Chen & Zou 1979CT83
    |    `--P. megasiphonatum Chen & Zou 1979CT83
    |--Sinolebetoceras Chen & Qi 1979CT83
    |    `--S. compressum Chen & Qi 1979CT83
    |--Cumberloceras Ulrich, Foerste & Miller 1943FG64
    |    |--*C. buttsi Ulrich, Foerste & Miller 1943FG64
    |    `--C. elwayense Ulrich, Foerste & Miller 1943P68
    |--Robsonoceras Ulrich & Foerste 1933FG64
    |    |--*R. robsonense (Walcott 1924) [=Ellesmeroceras robsonensis]FG64
    |    `--R. manitouense Ulrich, Foerste et al. 1944P68
    |--Clelandoceras Ulrich, Foerste et al. 1944FG64
    |    |--*C. elongatum Ulrich, Foerste et al. 1944FG64
    |    `--C. breve Ulrich, Foerste et al. 1944P68
    |--Woosteroceras Ulrich, Foerste et al. 1944FG64
    |    |--*W. trempealeauense Ulrich, Foerste et al. 1944FG64
    |    `--W. cherokeense Ulrich, Foerste et al. 1944P68
    |--Qiushugouceras Chen & Teichert 1983CT83
    |    |--*Q. inclinatum Chen & Teichert 1983CT83
    |    `--Q. magnum Chen & Teichert 1983CT83
    |--Tanycameroceras Chen & Qi 1979CT83
    |    |--*T. anhuiense Chen & Qi in Chen et al. 1979CT83
    |    `--T. amplum Chen & Qi in Chen et al. 1980CT83
    |--Shantungendoceras Sun 1937FG64
    |    |--*S. conicum Sun 1937FG64
    |    |--S. curvatum Sun 1937CT83
    |    `--S. nanshanense Sun 1937CT83
    |--Buehleroceras Ulrich, Foerste & Miller 1943FG64
    |    |--*B. compressum Ulrich, Foerste & Miller 1943FG64
    |    |--B. apertum Ulrich, Foerste & Miller 1943P68
    |    |--B. luthei Ulrich, Foerste & Miller 1943P68
    |    `--B. modestum Ulrich, Foerste & Miller 1943P68
    |--Albertoceras Ulrich & Foerste 1926FG64
    |    |--*A. walcotti Ulrich & Foerste 1926FG64
    |    |--A. gracillimum Ulrich, Foerste et al. 1944P68
    |    |--A. minimum Ulrich, Foerste et al. 1944P68
    |    `--A. staufferi Ulrich, Foerste et al. 1944P68
    |--Anhuiceras Zou & Chen in Chen et al. 1979CT83
    |    |--A. concavoseptum Zou & Chen 1979CT83
    |    |--A. elongatum Zou & Chen 1979CT83
    |    |--A. longicervicum Zou & Chen 1979CT83
    |    `--A. qingshuihense Zou & Chen 1979CT83
    |--Chabactoceras Malinovskaya 1964CT83
    |    |--*C. balaschovi Malinovskaya 1964CT83
    |    |--C. annulatum Chen & Qi 1982CT83
    |    |--C. planum Chen & Teichert 1983CT83
    |    `--C. sinicum Chen & Qi 1982CT83
    |--Eoclarkoceras Chen & Qi 1979CT83
    |    |--E. anhuiense Chen & Qi 1979CT83
    |    |--E. lunanense Chen & Qi 1980CT83
    |    |--E. parvum Chen & Qi 1980CT83
    |    `--E. subcurvatum (Kobayashi 1933) [=Ectenolites subcurvatus]CT83
    |--Caseoceras Ulrich, Foerste & Miller 1943FG64
    |    |--*C. contractum Ulrich, Foerste & Miller 1943FG64
    |    |--C. conicum Ulrich, Foerste & Miller 1943P68
    |    |--C. holtedahliP68
    |    |--C. intermedium Ulrich, Foerste & Miller 1943P68
    |    |--C. nitidum Ulrich, Foerste & Miller 1943P68
    |    |--C. sequatchiense Ulrich, Foerste & Miller 1943P68
    |    `--C. subconicum Ulrich, Foerste & Miller 1943P68
    |--Eburoceras Zou & Chen 1979CT83
    |    |--*E. jiagouense Zou & Chen in Chen et al. 1979CT83
    |    |--E. aduncum Chen & Qi 1980CT83
    |    |--E. grossotubulum Chen & Qi in Chen et al. 1980CT83
    |    |--E. hookiforme Zou & Chen 1979CT83
    |    |--E. pissinum Chen & Qi 1980CT83
    |    |--E. qianshanense Zou & Chen 1979CT83
    |    `--E. suxianense Zou & Chen 1979CT83
    |--Ellesmeroceras Foerste 1921 (see below for synonymy)FG64
    |    |--*E. scheii Foerste 1921 [=*Ellesmereoceras scheii]FG64
    |    |--E. abruptum Kobayashi 1933CT83
    |    |--E. bridgei Flower 1941P68
    |    |--E. curvatum Kobayashi 1931CT83
    |    |--E. elongatum Kobayashi 1931CT83
    |    |--E. expansum Flower 1941P68
    |    |--E. foerstei Kobayashi 1933CT83
    |    |--E. higdonense Ulrich, Foerste et al. 1944P68
    |    |--E. hotchkissi Ulrich, Foerste et al. 1944P68
    |    `--E. subcirculare Kobayashi 1931CT83
    |--Oneotoceras Ulrich 1926FG64
    |    |--*O. loculosum (Hall 1861) [=Cyrtoceras loculosum]FG64
    |    |--O. curvatum Ulrich, Foerste & Miller 1943P68
    |    |--O. divaricatum Ulrich, Foerste & Miller 1943P68
    |    |--O. extremum Ulrich, Foerste & Miller 1943P68
    |    |--O. husseyi Ulrich, Foerste & Miller 1943P68
    |    |--O. impressum Ulrich, Foerste & Miller 1943P68
    |    |--O. madisonense Ulrich, Foerste & Miller 1943P68
    |    |--O. magnum Ulrich, Foerste & Miller 1943P68
    |    |--O. percurvatum Ulrich, Foerste & Miller 1943P68
    |    `--O. wisconsinense Ulrich, Foerste & Miller 1943P68
    |--Conocerina Ulrich & Foerste 1936 [=Concerina (l. c.)]FG64
    |    |--*C. brevis Ulrich & Foerste 1936FG64
    |    |--C. arrecta Ulrich, Foerste & Miller 1943P68
    |    |--C. bassleri Ulrich, Foerste & Miller 1943P68
    |    |--C. beaniFG64
    |    |--C. cloudi Unklesbay 1954P68
    |    |--C. magna Ulrich, Foerste & Miller 1943P68
    |    |--C. missouriensis Ulrich, Foerste & Miller 1943P68
    |    |--C. obliqua Ulrich, Foerste & Miller 1943P68
    |    |--C. raaschiP68
    |    `--C. sublonga Ulrich, Foerste & Miller 1943P68
    |--Ectenolites Ulrich & Foerste 1936 [incl. Tamdoceras Malinovskaya 1964]CT83
    |    |--E. conicus (Malinovskaya 1964) [=Tamdoceras conicum]CT83
    |    |--E. curvatus (Kobayashi 1931)CT83
    |    |--E. flexuostriatum Kobayashi 1931CT83
    |    |--E. lisogorae (Malinovskaya 1964) [=Tamdoceras lisogorae]CT83
    |    |--E. longicameratus (Malinovskaya 1964) [=Tamdoceras longicameratum]CT83
    |    |--E. penecilin Flower 1964CT83
    |    |--E. petilus Chen & Qi 1982CT83
    |    |--E. primus Flower 1964CT83
    |    |--E. pusillus Chen & Teichert 1983CT83
    |    `--E. qiushugouensis Chen & Teichert 1983CT83
    |--Dakeoceras Ulrich & Foerste 1931FG64
    |    |--*D. normale Ulrich & Foerste 1931FG64
    |    |--D. corniculum Ulrich, Foerste & Miller 1943P68
    |    |--D. dilatatum Ulrich, Foerste & Miller 1943P68
    |    |--D. divergens Ulrich, Foerste & Miller 1943P68
    |    |--D. magnisiphonatum Ulrich, Foerste & Miller 1943P68
    |    |--D. perretrorsum Ulrich, Foerste & Miller 1943P68
    |    |--D. retrorsum Ulrich, Foerste & Miller 1943P68
    |    |--D. reversum Ulrich, Foerste & Miller 1943P68
    |    |--D. separatum Ulrich, Foerste & Miller 1943P68
    |    |--D. subcurvatum Ulrich & Foerste 1931P68
    |    |--D. undulatum Ulrich, Foerste & Miller 1943P68
    |    `--D. vernonense Ulrich, Foerste & Miller 1943P68
    |--Levisoceras Foerste 1925FG64
    |    |--*L. mercurius (Billings 1865) [=Cyrtocerina mercurius]FG64
    |    |--L. arctedorsatum Ulrich, Foerste & Miller 1943P68
    |    |--L. complanatum Ulrich, Foerste & Miller 1943P68
    |    |--L. constrictum Ulrich, Foerste & Miller 1943P68
    |    |--L. curvatum Ulrich, Foerste & Miller 1943P68
    |    |--L. ellipticum Ulrich, Foerste & Miller 1943P68
    |    |--L. instabile Ulrich, Foerste & Miller 1943P68
    |    |--L. magnum Unklesbay 1954P68
    |    |--L. obliquatum Ulrich, Foerste & Miller 1943P68
    |    |--L. percurvatum Ulrich, Foerste & Miller 1943P68
    |    |--L. raaschi Ulrich, Foerste & Miller 1943P68
    |    |--L. subgracile Ulrich, Foerste & Miller 1943P68
    |    `--L. transitum Ulrich, Foerste & Miller 1943P68
    |--Burenoceras Ulrich & Foerste 1931FG64
    |    |--*B. pumilum Ulrich & Foerste 1931FG64
    |    |--B. angulare Ulrich, Foerste & Miller 1943P68
    |    |--B. compressum Ulrich, Foerste & Miller 1943P68
    |    |--B. curticei Ulrich, Foerste & Miller 1943P68
    |    |--B. dickhauti Ulrich, Foerste & Miller 1943P68
    |    |--B. dilatatum Ulrich, Foerste & Miller 1943P68
    |    |--B. expandum Ulrich & Foerste 1931P68
    |    |--B. huzzahense Ulrich, Foerste & Miller 1943P68
    |    |--B. minor Ulrich, Foerste & Miller 1943P68
    |    |--B. oneotense Ulrich, Foerste & Miller 1943P68
    |    |--B. peramplum Ulrich, Foerste & Miller 1943P68
    |    |--B. percompressum Ulrich, Foerste & Miller 1943P68
    |    |--B. planiseptatum Ulrich, Foerste & Miller 1943P68
    |    |--B. reticulatum Kobayashi 1933CT83
    |    |--B. texanum Ulrich, Foerste & Miller 1943P68
    |    |--B. undosum Ulrich, Foerste & Miller 1943P68
    |    `--B. ungulatum Ulrich, Foerste & Miller 1943P68
    |--Eremoceras Hyatt 1884 [incl. Ectenoceras Ulrich & Foerste in Kobayashi 1933]FG64
    |    |--*E. syphax (Billings 1865) [=Cyrtoceras syphax]FG64
    |    |--‘Ectenoceras’ arbucklense Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ arcuosutum Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ chepultepecense Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ compressum Ulrich, Foerste & Miller 1943P68
    |    |--E. conoidale Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ ellipticum Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ exile Ulrich, Foerste & Miller 1943P68
    |    |--E. gasconadense Ulrich, Foerste & Miller 1943P68
    |    |--‘Cyrtoceras’ gracile Cleland 1903 [=Ectenoceras gracilis]P68
    |    |--‘Ectenoceras’ higdonense Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ laqueatum Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ longum Ulrich, Foerste & Miller 1943P68
    |    |--E. marylandicum Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ millsi Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ obliquum Ulrich, Foerste & Miller 1943P68
    |    |--E. perdilatum Ulrich, Foerste & Miller 1943P68
    |    |--E. pergracile (Ulrich, Foerste & Miller 1943)FG64, P68 [=Ectenoceras pergracileP68]
    |    |--‘Ectenoceras’ perobliquum Ulrich, Foerste & Miller 1943P68
    |    |--‘Ectenoceras’ ruedemanni Kobayashi 1933CT83
    |    |--E. sansabaense Unklesbay 1954P68
    |    |--E. subcurvatum (Kobayashi 1933)FG64, CT83 [=Ectenoceras subcurvatumCT83]
    |    |--E. subgracile (Ulrich & Foerste 1936)FG64, P68 [=Ectenolites subgracileP68]
    |    |--E. subrectum Ulrich, Foerste & Miller 1943P68
    |    |--E. teres Ulrich, Foerste & Miller 1943P68
    |    `--E. vernonense Ulrich, Foerste & Miller 1943P68
    `--Clarkoceras Ruedemann 1905 [=Clarkeoceras Flower 1941 non Wedekind 1918]FG64
         |--*C. newtonwinchelli (Clarke 1897) [=Piloceras newtonwinchelli, *Clarkeoceras newtonwinchelli]FG64
         |--C. affine Ulrich, Foerste & Miller 1943P68
         |--C. arcuatum Ulrich, Foerste & Miller 1943P68
         |--C. buttsi Ulrich, Foerste & Miller 1943P68
         |--C. calviniP68
         |--C. cinctum Ulrich, Foerste & Miller 1943P68
         |--C. confertum Ulrich, Foerste & Miller 1943P68
         |--C. crassum Ulrich, Foerste & Miller 1943P68
         |--C. cuneatum Ulrich, Foerste & Miller 1943P68
         |--C. curvatum Ulrich, Foerste & Miller 1943P68
         |--C. expansum Ulrich, Foerste & Miller 1943P68
         |--C. gracile Ulrich, Foerste & Miller 1943P68
         |--C. huzzahense Ulrich, Foerste & Miller 1943P68
         |--C. inconstans Ulrich, Foerste & Miller 1943P68
         |--C. jasperense Ulrich, Foerste & Miller 1943P68
         |--C. lawrencenseFG64
         |--C. lutheiFG64
         |--C. magnisiphonatum Ulrich, Foerste & Miller 1943P68
         |--C. minneiskense Ulrich, Foerste & Miller 1943P68
         |--C. modestum Ulrich, Foerste & Miller 1943P68
         |--C. multicameratum Ulrich, Foerste & Miller 1943P68
         |--C. parvisiphonatum Ulrich, Foerste & Miller 1943P68
         |--C. poulseni Kobayashi 1931CT83
         |--C. qianshanense Zou & Chen 1979CT83
         |--C. rectum Ulrich, Foerste & Miller 1943P68
         |--C. ruguliferum Ulrich, Foerste & Miller 1943P68
         |--C. subarcuatum Ulrich, Foerste & Miller 1943P68
         |--C. subcrassum Ulrich, Foerste & Miller 1943P68
         |--C. subrectum Ulrich, Foerste & Miller 1943P68
         |--C. vicinum Ulrich, Foerste & Miller 1943P68
         `--C. whitehallense Ulrich, Foerste & Miller 1943P68

Ellesmeroceras Foerste 1921 [=Ellesemroceras (l. c.), Ellesmereoceras Foerste 1924; incl. Metaellesmeroceras Flower 1956, Metallesmeroceras (l. c.)]^FG64

*Type species of generic name indicated

References

[CT83] Chen J.-Y. & C. Teichert. 1983. Cambrian Cephalopoda of China. Palaeontographica Abteilung A: Paläozoologie—Stratigraphie 181 (1–3): 1–102.

[EK12] Evans, D. H., & A. H. King. 2012. Resolving polyphyly within the Endocerida: the Bisonocerida nov., a new order of early Palaeozoic nautiloids. Geobios 45 (1): 19–28.

[FG64] Furnish, W. M., & B. F. Glenister. 1964. Nautiloidea—Ellesmerocerida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt K. Mollusca 3. Cephalopoda—General Features—Endoceratoidea—Actinoceratoidea—Nautiloidea—Bactritoidea pp. K129–K159. The Geological Society of America and the University of Kansas Press.

[K93] King, A. H. 1993. Mollusca: Cephalopoda (Nautiloidea). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 169–188. Chapman & Hall: London.

[KE19] King, A. H., & D. H. Evans. 2019. High-level classification of the nautiloid cephalopods: a proposal for the revision of the Treatise Part K. Swiss Journal of Palaeontology 138: 65–85.

[M13] Mutvei, H. 2013. Characterization of nautiloid orders Ellesmerocerida, Oncocerida, Tarphycerida, Discosorida and Ascocerida: new superorder Multiceratoidea. GFF 135 (2): 171–183.

[P68] Purnell, L. R. 1968. Catalog of the type specimens of invertebrate fossils. Part I: Paleozoic Cephalopoda. United States National Museum Bulletin 262: 1–198.