
Belongs within: Gelechioidea.
The Ethmiinae are a group of moths with larvae feeding on Boraginaceae. Infestation with Ethmia heliomela has been known to cause defoliation of its host Ehretia acuminata (Nielsen & Common 1991).
Characters (from Nielsen & Common 1991, as Ethmiidae): Small to medium sized; head smooth scaled; ocelli absent; antennae usually without pecten on scape; maxillary palps usually four-segmented, folded over base of proboscis; labial palps recurved; epiphysis present; hind tibiae with long filiform scales; fore wing without pterostigma, R4 and R5 stalked, R5 to costa or apex, CuP present at least towards margind; hind wing with Rs and M1 well separated at base, divergent, M2 not arising nearer to M3 than M1, CuP present; frenulum in female with three bristles; abdomen not spined; male genitalia with uncus often bifurcate or reduced, gnathos sometimes absent, valva usually deeply divided. Larva usually with some secondary setae at least in SV groups, crochets in bi- or uniordinal circle, mesoseries or mesal penellipse; usually beneath light webbing on Boraginaceae. Pupa with a pair of ventral, leg-like processes on abdominal segment 9, each with a series of hooked apical setae.
<==Ethmiinae [Ethmiidae] | i. s.: ‘Symmoca’ maculifera Matsumura 1931G63 |--Dasyethmia Danilevsky 1969K07 | `--*D. hiemalis Danilevsky 1969K07 `--EthmiaK07 |--E. afghana Sattler 1967K07 |--E. assamensis Butler 1879 [incl. Symmoca okinawana Matsumura 1931]G63 |--E. bipunctella (Fabricius 1775)K07 |--E. cirrhocnemia (Lederer 1870)K07 |--E. clytodoxaNC91 |--E. comitellaK07 | |--E. c. comitellaK07 | `--E. c. steppella Dubatolov & Ustjuzhanin 1997K07 |--E. defreinai Ganev 1984K07 |--E. derbendella Sattler 1967K07 |--E. distigmatellaK07 |--E. elimatella Danilevsky 1975 [=E. simatalla Danilevsky 1980 (n. n.)]K07 |--E. euphoria Kun 2007K07 |--E. hakkarica Koçak 1986K07 |--E. heliomelaNC91 |--E. hemadelphaNC91 |--E. infelix Meyrick 1914 [incl. E. kurdistanella Amsel 1959]K07 |--E. interposita Sattler 1967K07 |--E. iranella (Zerny 1940)K07 |--E. kabulica Amsel 1969K07 |--E. lecmimaK07 | |--E. l. lecmimaK07 | `--E. l. amsel Kemal & Koçak 2005K07 |--E. pagiopa Sattler 1967K07 |--E. percisa Kun 2007K07 |--E. quadrinotellaK07 | |--E. q. quadrinotellaK07 | `--E. q. heratella Amsel 1969K07 |--E. sattleri Kun 2007K07 |--E. sibirica Danilevsky 1975K07 |--E. similis Sattler 1967K07 |--E. soljanikovi Danilevsky & Zagulajev 1975K07 |--E. sphaerostichaNC91 |--E. suspecta Sattler 1967K07 |--E. thoraeaNC91 |--E. turkmeniella Dubatolov & Ustjuzhanin 1997K07 |--E. viduaK07 | |--E. v. viduaK07 | `--E. v. flavilaterella Danilevsky 1975K07 `--E. wurtseri Amsel 1956K07
*Type species of generic name indicated
References
[G63] Gozmány, L. A. 1963. The family Symmocidae and the description of new taxa mainly from the Near East (Lepidoptera). Acta Zoologica Academiae Scientiarum Hungaricae 9 (1–2): 67–134.
[K07] Kun, A. 2007. Studies on Palaearctic Ethmia Hübner, 1819 (Lepidoptera: Elachistidae). Annales Historico-Naturales Musei Nationalis Hungarici 99: 101–114.
[NC91] Nielsen, E. S., & I. F. B. Common. 1991. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 817–915. Melbourne University Press: Carlton (Victoria).