Belongs within: Perilampidae.
Contains: Eucharitinae.
The Eucharitidae are a group of wasps parasitic on ants. Like the closely related Perilampidae, eggs are not laid directly on the host; instead, they are laid amongst vegetation and hatch into mobile planidium larvae that then actively seek out a suitable host.
Synapomorphies (from Heraty et al. 2013): Axillae fused, forming a united transverse band such that scutellum is separated from trans-scutal articulation; transepimeral division absent; metascutellar arm less than half length of lateral panel but forming a convex flange and not reduced to a carina; propodeal spiracle circular; metapleural–propodeal supracoxal flange absent; uncus of fore wing absent; apex of protibial spur without a cleft; fore basitarsus without a comb of setae; petiole surface completely fused ventrally with no suture present; cercus flattened and button-like; ovipositor valvulae expanded.
<==Eucharitidae [Eucharidae, Eucharidinae]MH11 |--Akapala Girault 1934HB13, B88 [Akapalinae] | |--A. astriaticeps (Girault 1926) (see below for synonymy)B88 | `--A. rudis (Westwood 1874) [=Schizaspidia rudis]B88 `--GollumiellinaeHB13 |--Gollumiella Hedqvist 1978MH11, B88 | |--G. antennataHD09 | `--+--G. buffingtoniHD09 | |--G. minutaHD09 | `--+--*G. longipetiolataB88, HD09 | `--G. ochreataHD09 `--+--Anorasema Bouček 1988HB13, B88 | `--*A. pallidipes (Cameron 1909) [=Eucharis pallidipes]B88 `--+--EucharitinaeHB13 `--OraseminaeHB13 |--ParasemoraB88 `--Orasema Cameron 1884 (see below for synonymy)B88 | i. s.: *O. stramineipesB88 | O. assectatorB88 | O. gemma Girault 1932B88 | O. palgravei Girault 1922B88 | O. purpureoventris (Cameron 1909) (see below for synonymy)B88 | O. simplexMH11 | O. theocles (Walker 1839) (see below for synonymy)B88 | ‘*Eucharomorpha’ worcesteri Girault 1913B88 |--O. glabraMH11 |--+--O. synemporaMH11 | `--Orasemorpha Bouček 1988MH11, B88 [=Eucharomorpha Girault 1913 non Girault 1913B88] | |--*O. viridis (Girault 1913) [=*Eucharomorpha viridis]B88 | |--O. bidentata (Girault 1940) [=Eucharomorpha bidentata]B88 | |--O. dubia (Girault 1913) [=Eucharomorpha dubia]B88 | |--O. eribotes (Walker 1839) (see below for synonymy)B88 | |--O. fuscipes (Girault 1913) [=Eucharomorpha fuscipes]B88 | |--O. goethei (Girault 1934) [=Eucharomorpha goethei]B88 | |--O. partiglabra (Girault 1940) [=Eucharomorpha partiglabra]B88 | |--O. pyttalus (Walker 1846) [=Eucharis pyttalus]B88 | |--O. tridentata (Girault 1915) [=Eucharomorpha tridentata]B88 | |--O. varidentata (Girault 1936) [=Eucharomorpha varidentata]B88 | |--O. wheeleri (Brues 1934) [=Eucharomorpha wheeleri]B88 | `--O. xeniades (Walker 1839) [=Eucharis xeniades]B88 `--+--+--+--+--O. communisMH11 | | | `--O. valgius (Walker 1839)HD09, B88 (see below for synonymy) | | `--TimioderusHB13 | | |--T. acuminatusHB13 | | |--T. peridentatusMH11 | | `--T. refringensMH11 | `--Indosema indicaHB13 `--+--+--O. simulatrixHB13 | `--O. texanaMH11 `--+--+--O. coloradensisMH11 | `--O. robertsoniHD09 `--+--O. delicatula (Walker 1862)MH11, B88 [=Eucharis delicatulaB88, Psilogaster delicatulaB88] `--+--O. rapoHD09 `--+--O. costaricensisHD09 `--O. minutissimaHD09
Akapala astriaticeps (Girault 1926) [=Kapala astriaticeps; incl. *Akapala astriaticeps Girault 1934 non Kapala astriaticeps Girault 1926]B88
Orasema Cameron 1884 [incl. Eucharomorpha Girault 1913, Semora Cameron 1910, Semorata Strand 1947, Semorella Ghesquière 1946]B88
Orasema purpureoventris (Cameron 1909) [=Eucharis purpureoventris, Gollumiella purpureoventris; incl. Parapsilogaster laeviceps Gahan 1940, Losbanus laeviceps, Psilogaster nishidai Ishii & Nagasawa 1958, L. nishidai, L. peterseni Hedqvist 1978]B88
Orasema theocles (Walker 1839) [=Eucharis theocles, Psilogaster theocles; incl. Orasema emma Girault 1934]B88
Orasema valgius (Walker 1839)HD09, B88 [=Eucharis valgiusB88, Epimetagea valgiusB88, Parapsilogaster valgiusB88, Psilogasteroides valgiusB88; incl. Orasema pheidolophaga Girault 1913B88]
Orasemorpha eribotes (Walker 1839) [=Eucharis eribotes, Epimetagea eribotes, Psilogasteroides eribotes]B88
*Type species of generic name indicated
References
[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).
[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466–542.
[HD09] Heraty, J. M., & D. C. Darling. 2009. Fossil Eucharitidae and Perilampidae (Hymenoptera: Chalcidoidea). Zootaxa 2306: 1–16.
[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.