
Belongs within: Asteroideae.
Eupatorium is a Holarctic genus of perennial herbs bearing flowers in clusters of discoid heads. Some species are known as bonesets, in reference to the historical use of the common boneset E. perfoliatum to treat painful fevers.
Characters (Flora of North America): Perennials, 30–200 cm. Stems erect, usually not branched proximal to arrays of heads (from caudices or rhizomes). Leaves mostly cauline; usually opposite (rarely whorled, distal sometimes alternate); petiolate or sessile; blades usually 3-nerved from or distal to bases, or pinnately nerved, mostly deltate or ovate to lanceolate or linear (and intermediate shapes, sometimes elliptic, oblong, rhombic, or suborbiculate, sometimes pinnatifid, 1–2-pinnately, ternately, or palmately lobed), ultimate margins entire or toothed, faces glabrous or puberulent, pubescent, scabrous, or setulose, usually gland-dotted. Heads discoid, in corymbiform or diffuse to dense, paniculiform arrays. Involucres obconic to ellipsoid, 1–3(–5+) mm diam. Phyllaries persistent, 7–15+ in 2–3(–4+) series, (usually green) 2–3-nerved, or not notably nerved, or pinnately nerved, elliptic, lanceolate, oblong, or obovate, usually unequal, sometimes ± equal (margins scarious, hyaline, apices rounded to acute or acuminate sometimes mucronate, faces usually puberulent or villous, usually gland-dotted, rarely glabrous). Receptacles flat or convex, epaleate. Florets (3–)5(–15+); corollas usually white, rarely pinkish, throats funnelform to campanulate, lobes 5, triangular; styles: bases sometimes enlarged, usually puberulent (rarely glabrous), branches mostly filiform. Cypselae (brownish to black) prismatic, 5-ribbed, usually glabrous, usually gland-dotted; pappi persistent, of 20–50 (whitish) barbellulate bristles in 1 series. x = 10.
<==Eupatorium
|--E. cabaionumJ87
|--E. cannabinumLDB98
|--E. capillifoliumACW01
|--E. coelestinumS72
|--E. elphantusBM76
|--E. flavidulumJ87
|--E. glabrumBT87
|--E. glandulosumT-W89
|--E. glehniT03
|--E. hemiteropododumBT87
|--E. illitumJ87
|--E. microchaetumJ87
|--E. mikaniodesM83
|--E. nervosumJ87
|--E. odoratumG01
|--E. perfoliatumC13
|--E. polystachyumRJ11
|--E. porphyrocladiumJ87
|--E. puberulumJ87
|--E. purpureumC38
|--E. ripariumH97
|--E. rugosumEBS98
|--E. selleanumJ87
|--E. stigmaticumJ87
|--E. urbaniiJ87
`--E. urticaefoliumC13
*Type species of generic name indicated
References
[ACW01] Aguilar, H., C. C. Childers & W. C. Welbourn. 2001. Relative abundance and seasonal occurrence of mites in the family Tydeidae on citrus in Florida. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 376–380. CSIRO Publishing: Melbourne.
[BT87] Baker, E. W., & D. M. Tuttle. 1987. The false spider mites of Mexico (Tenuipalpidae: Acari). United States Department of Agriculture, Technical Bulletin 1706: 1–237.
[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.
[C13] Cosens, A. 1913. Insect galls. Canadian Entomologist 45 (11): 380–384.
[C38] Crawford, J. C. 1938. Some new or little known Thysanoptera. Proceedings of the Entomological Society of Washington 40 (2): 35–43.
[EBS98] Elliott, K. J., L. R. Boring & W. T. Swank. 1998. Changes in vegetation structure and diversity after grass-to-forest succession in a southern Appalachian watershed. American Midland Naturalist 140: 219–232.
[G01] Gupta, S. K. 2001. A conspectus of natural enemies of phytophagous mites and mites as potential biocontrol agents of agricultural pests in India. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 484–497. CSIRO Publishing: Melbourne.
[H97] Hamilton, A. G. 1897. On domatia in certain Australian and other plants. Proceedings of the Linnean Society of New South Wales 21 (4): 758–792, pl. 57.
[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.
[LDB98] Lenssen, J. P. M., G. E. ten Dolle & C. W. P. M. Blom. 1998. The effect of flooding on the recruitment of reed marsh and tall forb plant species. Plant Ecology 139: 13–23.
[M83] Myers, R. L. 1983. Site susceptibility to invasion by the exotic tree Melaleuca quinquenervia in southern Florida. Journal of Applied Ecology 20: 645–658.
[RJ11] Rising, J. D., A. Jaramillo, J. L. Copete, P. G. Ryan & S. C. Madge. 2011. Family Emberizidae (buntings and New World sparrows). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 428–683. Lynx Edicions: Barcelona.
[S72] Shields, O. 1972. Flower visitation records for butterflies (Lepidoptera). Pan-Pacific Entomologist 48 (3): 189–203.
[T-W89] Tenison-Woods, J. E. 1889. On the vegetation of Malaysia. Proceedings of the Linnean Society of New South Wales, series 2, 4 (1): 9–106, pls 1–9.
[T03] Tsurusaki, N. 2003. Phenology and biology of harvestmen in and near Sapporo, Hokkaido, Japan, with some taxonomical notes on Nelima suzukii n. sp. and allies (Arachnida: Opiliones). Acta Arachnologica 52: 5–24.