Belongs within: Castorimorpha.
Contains: Thomomys, Geomys.
The Geomyidae, pocket gophers, are a group of burrowing rodents found in North and Central America. They share the presence of externally opening, fur-lined cheek pouches with the related kangaroo rats and pocket mice of the Heteromyidae, but are distinguished by their adaptions for a fossorial lifestyle (Hafner & Hafner 1983).
<==Geomyidae
| i. s.: ‘Sorex’ mexicanus Kerr 1792HH83
| DikkomysB74
| Heterogeomys hispidusG69
| |--H. h. hispidusG69
| |--H. h. isthmicusG69
| |--H. h. tehuantepecusG69
| `--H. h. torridusG69
| GregorymysMJ11
|--EntoptychusMJ11 [EntoptychinaeHH83]
|--PleurolicusMJ11 [PleurolicinaeHH83]
| `--P. sulcifronsTS96
`--GeomyinaeHH83
|--ParapliosaccomysHH83
`--+--ThomomysGE05
`--+--+--‘Thomomys’ bulbivorusFS15
| `--+--‘Thomomys’ townsendiiFS15
| `--+--‘Thomomys’ bottaeFS15
| | |--T. b. bottaeBT40
| | |--T. b. actuosusHHP83
| | |--T. b. analogus Goldman 1938MB86
| | |--T. b. anitaeG91
| | |--T. b. basilicae Benson & Tillotson 1940 (see below for synonymy)BT40
| | |--T. b. cultellusHHP83
| | |--T. b. internatusHHP83
| | |--T. b. jacinteusG91
| | |--T. b. occipitalis Dice 1925BT40
| | `--T. b. rubidusHHP83
| `--‘Thomomys’ umbrinusFS15
| |--T. u. umbinusMB86
| |--T. u. arriagensis Dalquest 1951MB86
| |--T. u. crassidens Nelson & Goldman 1934MB86
| |--T. u. durangae Nelson & Goldman 1934MB86
| |--T. u. goldmani Merriam 1901MB86
| |--T. u. musculus Nelson & Goldman 1934MB86
| |--T. u. newmaniMB86
| |--T. u. sheldoni Bailey 1915MB86
| `--T. u. zacatecae Nelson & Goldman 1934 [incl. T. u. enixus Nelson & Goldman 1934]MB86
`--+--GeomysFS15
`--+--Zygogeomys trichopusFS15
`--+--+--+--‘Orthogeomys’ hispidusGE05
| | `--‘Orthogeomys’ laniusFS15
| `--+--‘Orthogeomys’ underwoodiFS15
| `--+--‘Orthogeomys’ cavatorFS15
| `--+--‘Orthogeomys’ cherrieiFS15
| `--+--‘Orthogeomys’ matagalpaeFS15
| `--+--‘Orthogeomys’ dariensisFS15
| `--+--Macrogeomys heterodusHH83 [=O. (Macrogeomys) heterodusHH83, FS15]
| `--‘Orthogeomys’ thaeleriFS15
`--+--OrthogeomysFS15
| |--O. cuniculusG69
| `--O. (Orthogeomys) grandisHH83
| |--O. g. grandisG69
| |--O. g. alleniG69
| |--O. g. alvarezi Schaldach 1966G69
| |--O. g. carboG69
| |--O. g. felipensisG69
| |--O. g. nelsoniG69
| `--O. g. scalopsG69
`--+--CratogeomysFS15
| |--+--C. fumosusFS15
| | `--C. planicepsFS15
| `--+--+--C. castanopsFS15
| | `--C. goldmaniFS15
| `--+--C. perotensisFS15
| `--+--C. fulvescensFS15
| `--C. merriamiFS15
`--PappogeomysGE05
|--P. alcorni Russell 1957R57
|--P. bulleriGE05
|--P. castanopsMB86
| |--P. c. castanopsMB86
| |--P. c. goldmani (Merriam 1895)MB86
| |--P. c. rubellus (Nelson & Goldman 1934)MB86
| |--P. c. subnubilus (Nelson & Goldman 1934)MB86
| `--P. c. surculus Russell 1968MB86
|--P. fumosusIT07
|--P. gymnurusHH83
|--P. merriamiIT07
|--P. neglectusIT07
|--P. tylorhinusIT07
`--P. zinseriIT07
Thomomys bottae basilicae Benson & Tillotson 1940 [=T. bottae occipitalis Benson & Tillotson 1939 non Dice 1925]BT40
*Type species of generic name indicated
References
[BT40] Benson, S. B., & D. F. Tillotson. 1940. A substitute name for Thomomys bottae occipitalis Benson and Tillotson. Proceedings of the Biological Society of Washington 53: 93.
[B74] Bugge, J. 1974. The cephalic arterial system in insectivores, primates, rodents and lagomorphs, with special reference to the systematic classification. Acta Anatomica 87 (Suppl 62): 1–160.
[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.
[G69] Goodwin, G. G. 1969. Mammals from the State of Oaxaca, Mexico, in the American Museum of Natural History. Bulletin of the American Museum of Natural History 141 (1): 1–269, 40 pls.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[G91] Groves, C. P. 1991. A Theory of Human and Primate Evolution revised ed. Clarendon Press: Oxford.
[HH83] Hafner, J. C., & M. S. Hafner. 1983. Evolutionary relationships of heteromyid rodents. In Biology of Desert Rodents. Great Basin Naturalist Memoirs 7: 3–29.
[HHP83] Hafner, J. C., D. J. Hafner, J. L. Patton & M. F. Smith. 1983. Contact zones and the genetics of differentiation in the pocket gopher Thomomys bottae (Rodentia: Geomyidae). Systematic Zoology 32 (1): 1–20.
[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.
[MB86] Matson, J. O. & R. H. Baker. 1986. Mammals of Zacatecas. Special Publications, Museum of Texas Tech University 24: 1–88.
[MJ11] Meredith, R. W., J. E. Janečka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer & W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334: 521–524.
[R57] Russell, R. J. 1957. A new species of pocket gopher (genus Pappogeomys) from Jalisco, México. University of Kansas Publications, Museum of Natural History 9 (11): 357–361.
[TS96] Tedford, R. H., J. B. Swinehart, C. C. Swisher III, D. R. Prothero, S. A. King & T. E. Tierney. 1996. The Whitneyan-Arikareean transition in the High Plains. In: Prothero, D. R., & R. J. Emry (eds) The Terrestrial Eocene–Oligocene Transition in North America pp. 312–334. Cambridge University Press.