Belongs within: Harpalitae.
Gnathaphanus is a genus of ground beetles found in Asia and Australasia, characterised by conspicuous dorsal punctures on the elytra, and sponge-like soles of adhesive setae on the fore and mid tarsi of males.
Gnathaphanus Macleay 1825 [incl. Mirosarus Bates 1878, Pachauchenius Macleay 1864]LM87
|--*G. vulneripennis (Macleay 1825)LM87 [=Trechus (*Gnathaphanus) vulneripennisLM87; incl. Harpalus subcostatusB89]
|--G. aridus Blackburn 1892LM87
|--G. dispellens (Walker 1859) [=Harpalus dispellens, Anisodactylus dispellens]B89
|--G. froggatti (Macleay 1888) [=Diaphoromerus froggatti]LM87
|--G. herbaceus Sloane 1900LM87
|--G. latus Sloane 1900LM87
|--G. licinoides Hope 1842 [incl. Harpalus alternans Castelnau 1867, Catadromus impressus Montrouzier 1860]LM87
|--G. melbournensis (Castelnau 1867) (see below for synonymy)LM87
|--G. multipunctatus (Macleay 1888) (see below for synonymy)LM87
|--G. philippensis (Chevrolat 1841) [=Amblygnathus philippensis; incl. *Pachauchenius laeviceps Macleay 1864]LM87
|--G. picipes (Macleay 1864) [=Harpalus picipes; incl. Diaphoromerus sulcatulus Macleay 1888]LM87
|--G. porcatulus (Macleay 1888) [=Diaphoromerus porcatulus]LM87
|--G. pulcher (Dejean 1829) [=Harpalus pulcher]LM87
| |--G. p. pulcher [incl. Harpalus interstitialis Macleay 1864]LM87
| `--G. p. speciosus Schauberger 1935LM87
|--G. punctifer (Castelnau 1867) [=Harpalus punctiferus; incl. H. montanus Castelnau 1867]LM87
|--G. rectangulus Chaudoir 1878LM87
|--G. riverinae Sloane 1895LM87
|--G. sexpunctatus (Macleay 1888) [=Diaphoromerus sexpunctatus]LM87
`--G. whitei Sloane 1917LM87
Gnathaphanus melbournensis (Castelnau 1867) [=Harpalus melbournensis; incl. H. adelaidae Castelnau 1867, H. aeneonitens Macleay 1871, H. angustatus Macleay 1871, H. atroviridis Macleay 1871, H. gayndahensis Macleay 1871, *Mirosarus insularis Bates 1878, H. marginicollis Castelnau 1867, H. paroensis Castelnau 1867, H. planipennis Macleay 1871]LM87
Gnathaphanus multipunctatus (Macleay 1888) [=Diaphoromerus multipunctatus; incl. G. darwini Blackburn 1888, Harpalus impressipennis Castelnau 1867 non Dejean 1829, Dioryche upolensis Csiki 1915]LM87
*Type species of generic name indicated
References
[B89] Bates, H.-W. 1889. Contributions a la faune Indo-Chinoise. 3e mémoire. Carabidae. Annales de la Société Entomologique de France, 6e série 9: 261–286.
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.