Belongs within: Ostropales.
The Gomphillaceae are a family of mostly foliicolous fungi that are lichenised with green algae (Kirk et al. 2001).
Characters (Kirk et al. 2001): Thallus crustose. Ascomata apothecial, usually sessile, discoid, without well-defined margin. Interascal tissue of branched and anastomosing filiform paraphyses. Asci cylindrical, with only slight apical thickening, 1- to 8-spored. Ascospores ellipsoidal to filiform, with transverse septa, sometimes muriform. Anamorphs coelomycetous, spine- or brush-like, or peltate.
<==Gomphillaceae
|--Actinoplaca Müll. Arg. 1891 [=Actinoplacomyces Cif. & Tomas. 1953]KC01
|--Aulaxina Fée 1825KC01
|--Bullatina Vězda & Poelt 1987KC01
|--Calenia Müll. Arg. 1890 (see below for synonymy)KC01
|--Caleniopsis Vězda & Poelt 1987KC01
|--Gyalideopsis Vězda 1972 (nom. cons.) [incl. Diploschistella Vain. 1926 (nom. rej.)]KC01
|--Hippocrepidea Sérus. 1997KC01
|--Psorotheciopsis Rehm 1900EB03, KC01 (see below for synonymy)
|--Sagiolechia Massal. 1854 [incl. Rhexophiale Fr. 1860]KC01
|--Paratricharia Lücking 1997KC01
|--Gyalectidium Müll. Arg. 1881 (see below for synonymy)KC01
| `--G. filicinum [incl. *Tauromyces catenatulus]E99
|--Tricharia Fée 1825 (see below for synonymy)KC01
| |--T. couepiae (Bat.) Lücking & Sérus. 1998 [=*Aderkomyces couepiae]E99
| |--T. heterella [incl. *Psathyromyces rosacearum]E99
| `--‘*Microxyphiomyces’ manaosensisE99
|--Echinoplaca Fée 1824 (see below for synonymy)KC01
| |--E. amapensisE99
| |--E. leucotrichoides [incl. *Sporocybomyces pulcher]E99
| |--‘Phallomyces’ palmaeE99
| `--E. strigulaceaJK06
`--Gomphillus Nylander 1855F33 [incl. Baeopodium Trevis. 1857KC01, Mycetodium Massee 1856F33]
`--G. calicioides (Delise) Nylander 1854 (see below for synonymy)F33
|--G. c. f. calicioidesF33
|--G. c. f. microcephalus Nylander 1854F33
`--G. c. f. polycephalus Nylander 1854F33
Calenia Müll. Arg. 1890 [=Caleniomyces Cif. & Tomas. 1953; incl. Phlyctidium Müll. Arg. 1888 nec Wallr. 1833 nec (Braun) Rabenh. 1868]KC01
Echinoplaca Fée 1824 [incl. Arthellopsis Vain. 1896, Arthotheliopsidomyces Cif. & Tomas. 1953, Arthotheliopsis Vain. 1896, Phallomyces Bat. & Valle 1961, Pycnodermellina Bat. & Maia 1957, Sporocybomyces Maia 1967]KC01
Gomphillus calicioides (Delise) Nylander 1854 [=Baeomyces calicioides Delise in Duby 1830, Berengeria calicioides (Delise) Massee 1855, Mycetodium calicioides (Delise) Massee 1856]F33
Gyalectidium Müll. Arg. 1881 [incl. Gonothecis Clem. 1909, Gonothecium (Vain.) Clem. & Shear 1931, Lopadiopsidomyces Cif. & Tomas. 1953, Lopadiopsis Vain. 1896, Tauromyces Cavalc. & Silva 1972]KC01
Psorotheciopsis Rehm 1900EB03, KC01 [incl. Conicosolen Schill. 1927KC01, Linhartia Sacc. & Syd. 1902KC01, Monospermella Speg. 1924KC01]
Tricharia Fée 1825 [incl. Aderkomyces Bat. 1961, Microxyphiomyces Bat., Valle & Peres 1961, Psathyromyces Bat. & Peres 1964]KC01
*Type species of generic name indicated
References
[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440–2755. Myconet 2: 1–41.
[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota—2003. Myconet 9: 1–89.
[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4 pt 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig.
[JK06] James, T. Y., F. Kauff, C. L. Schoch, P. B. Matheny, V. Hofstetter, C. J. Cox, G. Celio, C. Gueidan, E. Fraker, J. Miadlikowska, H. T. Lumbsch, A. Rauhut, V. Reeb, A. E. Arnold, A. Amtoft, J. E. Stajich, K. Hosaka, G.-H. Sung, D. Johnson, B. O’Rourke, M. Crockett, M. Binder, J. M. Curtis, J. C. Slot, Z. Wang, A. W. Wilson, A. Schüßler, J. E. Longcore, K. O’Donnell, S. Mozley-Standridge, D. Porter, P. M. Letcher, M. J. Powell, J. W. Taylor, M. M. White, G. W. Griffith, D. R. Davies, R. A. Humber, J. B. Morton, J. Sugiyama, A. Y. Rossman, J. D. Rogers, D. H. Pfister, D. Hewitt, K. Hansen, S. Hambleton, R. A. Shoemaker, J. Kohlmeyer, B. Volkmann-Kohlmeyer, R. A. Spotts, M. Serdani, P. W. Crous, K. W. Hughes, K. Matsuura, E. Langer, G. Langer, W. A. Untereiner, R. Lücking, B. Büdel, D. M. Geiser, A. Aptroot, P. Diederich, I. Schmitt, M. Schultz, R. Yahr, D. S. Hibbett, F. Lutzoni, D. J. McLaughlin, J. W. Spatafora & R. Vilgalys. 2006. Reconstructing the early evolution of Fungi using a six-gene phylogeny. Nature 443: 818–822.
[KC01] Kirk, P. M., P. F. Cannon, J. C. David & J. A. Stalpers. 2001. Ainsworth & Bisby’s Dictionary of the Fungi 9th ed. CAB International: Wallingford (UK).