Belongs within: Schizophora.
Contains: Anthomyzidae.
The Heleomyzidae are a group of flies characterised by strong, convergent postocellar bristles, wing with a subcostal but no humeral break, and a preapical dorsal bristle on each tibia. The circumscription of the group has been disputed; as recognised below, it may be paraphyletic to the family Anthomyzidae (Wiegmann et al. 2011) and some authors have divided it between a number of families.
Members of the tribe Rhinotorini have unusually tall heads with an excavated vertex and lacking postocellar bristles, and wings without spaced costal spines (McAlpine & Woodley 2010). The Trixoscelidini have ocellar bristles arising outside the ocellar triangle, alongside or in front of the anterior ocellus. Suillia has short orbital plates that slope mesad away from the eye margin and bear only a single strong orbital bristle (McAlpine & Woodley 2010).
<==Heleomyzidae [Cnemospathidae, Helomyzidae]
|--Trixoscelidini [Trixoscelidae, Trixoscelididae]MW10
| |--Spilochroa ornataMW10
| |--Zagonia flavicornisT87
| |--ParatrixoscelisT87
| |--PsiloplogiaT87
| `--TrixoscelisWT11
| |--T. frontalisT87
| `--T. fumipennisMW10
`--+--AnthomyzidaeWT11
`--Suillia [Suilliinae, Suilliini]WT11
|--S. iniensMW10
|--S. longipennisMW10
|--S. luridaGP87
|--S. punctulataMW10
`--S. variegataWT11
Heleomyzidae incertae sedis:
ChiropteromyzaGE05
Heleomyzini [Heleomyzinae]MW10
|--Tephrochlamys rufiventrisMW10, GP87
|--Oecothea [incl. Aecothea, Neoecothea]MW10
| `--O. fenestralis [incl. Aecothea fidelis]GP87
|--Scoliocentra [incl. Amoebaleria]MW10
| `--S. helvola [=Amoebaleria helvola]MW10
|--PseudoleriaMW10
| |--P. pectinataCM70
| `--P. placataMW10 [incl. P. crassataGP87]
`--HeleomyzaGP87
|--H. maculipennis [=Scoliocentra (Leriola) maculipennis; incl. H. difficilis]GP87
|--H. serrataGP87
`--H. tigrinaL49
Cinderella [Cinderellini]MW10
`--C. lampraMW10
Borboropsis [Borboropsidae]GP87
`--B. puberula [incl. B. fulviceps]GP87
Oldenbergiella brumalisGP87
LutomyiaGP87
|--L. hemipteraGP87
`--L. spurcaGP87
Rhinotorini [Rhinotoridae]MW10
|--RhinotoraMW10
|--RhinotoroidesM87
|--AnastomyzaM87
|--ApophoneuraM87
|--ZentulaM87
|--NeorhinotoraMW10
| |--N. aristalisMW10
| `--N. diversaMW10
`--CairnsimyiaM87
|--C. aroanaO98
|--C. excavataM87
`--C. robustaM87
Paraneossos [Gephyromyzini]MW10
`--P. arizonicusGP87
Prosopantrum [Cnemospathidini]MW10
`--P. flavifronsMW10
Tapeigaster [Tapeigastrini]CM91
`--T. annulipesCM91
AllophylopsiniCM91
|--Diplogeomyza diaphoraCM91
`--AllophylopsisCM70
NeossosT87
AnorostomaGP87
|--A. coloradenseGP87
`--A. curraniO98
Porsenus johnsoniGP87
Allophyla atricornis [incl. A. laevis]GP87
Morpholeria tristisGP87
SchroederellaGP87
|--S. inersGP87
`--S. luteoalaGP87
OrbelliaGP87
EccoptomeraGP87
NeoleriaGP87
SpanopareaGP87
AnypotactaGP87
Trichochlamys borealisGP87
CephodapedonT87
Chaetohelomyza electricaRJ93
ElectroleriaP92
ProtosuilliaP92
*Type species of generic name indicated
References
[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.
[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).
[GP87] Gill, G. D., & B. V. Peterson. 1987. Heleomyzidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 973–980. Research Branch, Agriculture Canada.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.
[M87] McAlpine, J. F. 1987. Rhinotoridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 989–992. Research Branch, Agriculture Canada.
[MW10] McAlpine, D. K., & N. E. Woodley. 2010. Heleomyzidae (heleomyzid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1159–1164. NRC Research Press: Ottawa.
[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.
[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
[T87] Teskey, H. J. 1987. Trixoscelididae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 981–984. Research Branch, Agriculture Canada.
[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.