Belongs within: Psocomorpha.
Contains: Elipsocidae, Lachesilloidea, Pseudocaecilioidea, Peripsocoidea.
The Homilopsocidea are a clade of bark-lice supported by two synapomorphies: the presence of an egg-guide extending from the dorsodistal margin of the female subgenital plate, and dorsal swelling of the dorsal valve of the gonapophyses (Yoshizawa 2002). The Lesneiidae are a family of uncertain affinities within this clade in which females have fused abdominal segments and lack gonapophyses (Schmidt & New 2004).
Characters (Yoshizawa 2002): Head usually with rounded vertex except sharply pointed in Calopsocidae; postclypeus usually well bulged; epistomal suture with or without well-developed internal ridge; anteclypeus variable; male eyes usually much larger than female’s; ocelli complete, usually clustered on weakly developed tubercle; mandible short, with rounded outer margin except Calopsocidae with somewhat angled outer margin; lacinial tip usually bicuspid; galea usually flattened; labial palpus rounded. Prothorax less bulged dorsally; pterothorax strongly bulged dorsally; mesothorax with broad precoxal bridge and narrow trochantin; metaepisternum with broad membranous region. Forewing with various coloration and markings; posteroproximal corner smoothly rounded; venation generally of Caecilius-type. Hindwing with rounded posteroproximal corner; Sc faint. Male genitalia variable; phallosome complete. Female genitalia: subgenital plate usually with distal projection extended from dorsal margin. Gonapophyses usually complete; dorsal valve swollen dorsally, usually with subapical ventral process.
<==Homilopsocidea [Calopsocoidea, Elipsocoidea, Reuterellidae]
| i. s.: Lesneiidae [Lesneiinae]SN04
| |--Hemicaecilius Enderlein 1903SN04
| `--Lesneia Badonnel 1931SN04
| |--*L. capensis Badonnel 1931SN04
| |--L. niger Broadhead & Richards 1982SN04
| |--L. pulchra Broadhead & Richards 1982SN04
| `--L. stuvkenbergi Badonnel 1963SN04
|--+--ElipsocidaeY02
| `--Drymopsocus Smithers 1963SN04
| `--*D. brunneus Smithers 1963SN04
`--+--LachesilloideaY02
`--+--PseudocaecilioideaY02
|--PeripsocoideaY02
`--SabulopsocidaeSN04
|--Sabulopsocus Smithers 1969SN04
| `--*S. tractuosus Smithers 1969SN04
`--Moapsocus Schmidt & New 2004SN04
`--*M. angelicus Schmidt & New 2004SN04
*Type species of generic name indicated
References
[SN04] Schmidt, E. R., & T. R. New. 2004. A systematic and phylogenetic revision of the family Elipsocidae (Insecta: Psocoptera), with the erection of two new families: Lesneiidae and Sabulopsocidae. Invertebrate Systematics 18: 157–213.
[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.