Belongs within: Sordariomyceta.
The Hyaloscyphaceae are a group of fungi saprobic on wood and herbaceous material (Kirk et al. 2001).
Characters (Kirk et al. 2001): Stromata absent. Ascomata usually small, flat or concave, excipulum soft, fleshy, usually composed of prismatic or isodiametric cells, almost always with conspicuous, sometimes ornamented hairs surrounding disc. Interascal tissue of simple paraphyses, sometimes lanceolate. Asci small, with J+ or J- ring. Ascospores small, usually elongated, sometimes septate. Anamorphs rarely noted, when so hyphomycetous.
<==Hyaloscyphaceae
|--Hyaloscypha Boud. 1885 (see below for synonymy)KC01
| `--H. monodictysHM17
|--Lachnum Retz. 1779 (see below for synonymy)KC01
| |--L. bicolorLK04
| `--L. virgineumJK06
`--Arachnopezizoideae [Arachnopezizeae]KC01
|--Arachnopeziza Fuckel 1870 [incl. Arachnopezizella Kirschst. 1938, Arachnoscypha Boud. 1885]KC01
| `--A. aureliaE99
`--Pubigera Baral, Gminder & Svrcek 1995E99
`--*P. subvillosa (Rehm) Baral, Gminder & Svrcek 1995E99
Hyaloscyphaceae incertae sedis:
Amicodisca Svrček 1987KC01
Antinoa Velen 1934EB03O, KC01
Asperopilum Spooner 1987KC01
Austropezia Spooner 1987KC01
Betulina Velen. 1947KC01
Calycellina Höhn. 1918 [incl. Scutoscypha Graddon 1980]KC01
Calycina Nees ex Gray 1821EB03O, KC01 (see below for synonymy)
Calyptellopsis Svrček 1986KC01
Chrysothallus Velen. 1934KC01
Ciliolarina Svrček 1977KC01
Ciliosculum Kirschst. 1941KC01
Cistellina Raitv. 1978KC01
Clavidisculum Kirschst. 1938KC01
Dasyscyphella Tranzschel 1898KC01
Dematioscypha Svrček 1977 (see below for synonymy)KC01
Dimorphotricha Spooner 1987KC01
Echinula Graddon 1977KC01
Eriopezia (Sacc.) Rehm 1892KC01
Fuscolachnum Haines 1989KC01
Fuscoscypha Svrček 1987KC01
Graddonidiscus Raitv. & Galán 1992KC01
Hamatocanthoscypha Svrček 1977 (see below for synonymy)KC01
Hegermila Raitv. 1995KC01
Hyalacrotes (Korf & Kohn) Raitv. 1991KC01
Hyalopeziza Fuckel 1870 [incl. Hyalotricha Dennis 1949, Pseudolachnea Velen. 1934 non Ranoj. 1910]KC01
Hydrocina Scheuer 1991KC01
Hyphodiscus Kirschst. 1906 [incl. Incrupilella Svrček 1986]KC01
Incrucipulum Baral 1985KC01
Incrupila Raitv. 1970KC01
Lachnellula Karst. 1884 [incl. Trichoscypha Boud. 1885 non Hook. f. 1862, Trichoscyphella Nannf. 1932]KC01
Lasiobelonium Ellis & Everh. 1897 [incl. Lasiobelonis Clem. & Shear 1931, Sponheimeria Kirschst. 1941]KC01
Microscypha Syd. & Syd. 1919 [incl. Mollisiaster Kirschst. 1939]KC01
Mollisina Höhn. ex Weese 1926 [incl. Dendrotrichoscypha Svrček 1977]KC01
Otwaya Beaton 1978KC01
Parachnopeziza Korf 1978KC01
Perrotia Boud. 1901KC01
Phaeoscypha Spooner 1984KC01
Pithyella Boud. 1885 [incl. Helotiopsis Höhn. 1910]KC01
Polaroscyphus Huhtinen 1987KC01
Proliferodiscus Haines & Dumont 1983 [incl. Farinodiscus Svrček 1987]KC01
Proprioscypha Spooner 1987KC01
Protounguicularia Raitv. & Galán 1986KC01
Psilachnum Höhn. 1926KC01
Psilocistella Svrček 1977KC01
Rodwayella Spooner 1986KC01
Solenopezia Sacc. 1889 [incl. Niveostoma Svrček 1988]KC01
Tapesina Lambotte 1887KC01
`--*T. griseovitellina [incl. Chalara rubi]EB03N
Trichopeziza Fuckel 1870KC01
Unguicularia Höhn. 1905 (see below for synonymy)KC01
Unguiculariella Thind & Sharma 1990KC01
Unguiculella Höhn. 1906 (see below for synonymy)KC01
Urceolella Boud. 1885 [incl. Pilatia Velen. 1934]KC01
Velutaria Fuckel 1870 [incl. Schweinitzia Massee 1895 nec Elliott ex Nuttall 1818 nec Grev. 1823]KC01
Venturiocistella Raitv. 1978KC01
Albotricha Raitv. 1970KC01
Cistella Quél. 1886 (nom. cons.) nec Gistl 1848 (ICZN) nec Gray 1853 (ICZN) [=Discocistella Svrček 1962]KC01
Olla Velen. 1934 [incl. Pseudoolla Velen. 1934]KC01
Phialina Höhn. 1926 (see below for synonymy)KC01
Polydesmia Boud. 1885KC01
Calycina Nees ex Gray 1821EB03O, KC01 [incl. Calycella (Fr.) Boud. 1885KC01, Cystopezizella Svrček 1983KC01, Eubelonis Clem. 1909KC01, Pezizella Fuckel 1870KC01]
Dematioscypha Svrček 1977 [incl. Goidanichia Arnaud 1954 non Tomas. & Cif. 1952, Goidanichiella Arnaud ex Barron 1968, Goidanichiomyces Cif. & Tomas. 1953, Haplographium Berk. & Broome 1859, Schizocephalum Preuss 1852]KC01
Hamatocanthoscypha Svrček 1977 [=Uncinia Velen. 1934 non Pers. 1807, Unciniella Holm & Holm 1977; incl. Debaryoscyphus Arendh. & Sharma 1986]KC01
Hyaloscypha Boud. 1885 [incl. Cheiromycella Höhn. 1910, Clathrosphaerina Beverw. 1951, Eupezizella Höhn. 1926, Pseudaegerita Crane & Schokn. 1981, Truncicola Velen. 1934; Hyaloscyphoideae]KC01
Lachnum Retz. 1779 [incl. Arenaea Penz. & Sacc. 1901, Belonidium Mont. & Durieu 1846, Brunnipila Baral 1985, Capitotricha (Raitv.) Baral 1985, Chaetoscypha Syd. 1924, Dasypezis Clem. 1909, Dasyscypha Fuckel 1870, Dasyscyphus Gray 1821, Dyslachnum Clem. 1909, Erinella Quél. 1886, Erinellina Seaver 1951, Erioscypha Kirschst. 1938, Erioscyphella Kirschst. 1938, Helolachnum Torrend 1910, Hyphoscypha Bres. 1903, Lachnaster Höhn. 1917, Lachnella Boud. 1885 non Fr. 1836, Lachnobelonium Höhn. 1926, Lasiobelonium (Sacc.) Sacc. & Syd. 1899 non Ellis & Everh. 1897, Neodasyscypha Spooner 1987 (nom. inv.), Pezizellaster Höhn. 1917, Trichopezizella Dennis ex Raitv. 1969; Lachnoideae]KC01
Phialina Höhn. 1926 nec Bory de St. Vincent 1824 (ICZN) nec Costa 1856 (ICZN) [incl. Phialoscypha Raitv. 1977, Setoscypha Velen. 1934]KC01
Unguicularia Höhn. 1905 [incl. Catinella Kirschst. 1924 nec Pease 1870 (ICZN) nec Stache 1877 (ICZN) nec Boud. 1907, Chytrella Kirschst. 1941, Mycopandora Velen. 1947, Phalothrix Clem. 1909]KC01
Unguiculella Höhn. 1906 [incl. Globulina Velen. 1934 nec Link 1820 nec d’Orbigny in De la Sagra 1839 (ICZN) nec Speg. 1888]KC01
*Type species of generic name indicated
References
[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440–2755. Myconet 2: 1–41.
[EB03N] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, T. Laessøe & G. Rambold (eds.) 2003. Notes on ascomycete systematics. Nos 3580–3623. Myconet 9: 91–103.
[EB03O] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota—2003. Myconet 9: 1–89.
[HM17] Hongsanan, S., S. S. N. Maharachchikumbura, K. D. Hyde, M. C. Samarakoon, R. Jeewon, Q. Zhao, A. M. Al-Sadi & A. H. Bahkali. 2017. An updated phylogeny of Sordariomycetes based on phylogenetic and molecular clock evidence. Fungal Diversity 84: 25–41.
[JK06] James, T. Y., F. Kauff, C. L. Schoch, P. B. Matheny, V. Hofstetter, C. J. Cox, G. Celio, C. Gueidan, E. Fraker, J. Miadlikowska, H. T. Lumbsch, A. Rauhut, V. Reeb, A. E. Arnold, A. Amtoft, J. E. Stajich, K. Hosaka, G.-H. Sung, D. Johnson, B. O’Rourke, M. Crockett, M. Binder, J. M. Curtis, J. C. Slot, Z. Wang, A. W. Wilson, A. Schüßler, J. E. Longcore, K. O’Donnell, S. Mozley-Standridge, D. Porter, P. M. Letcher, M. J. Powell, J. W. Taylor, M. M. White, G. W. Griffith, D. R. Davies, R. A. Humber, J. B. Morton, J. Sugiyama, A. Y. Rossman, J. D. Rogers, D. H. Pfister, D. Hewitt, K. Hansen, S. Hambleton, R. A. Shoemaker, J. Kohlmeyer, B. Volkmann-Kohlmeyer, R. A. Spotts, M. Serdani, P. W. Crous, K. W. Hughes, K. Matsuura, E. Langer, G. Langer, W. A. Untereiner, R. Lücking, B. Büdel, D. M. Geiser, A. Aptroot, P. Diederich, I. Schmitt, M. Schultz, R. Yahr, D. S. Hibbett, F. Lutzoni, D. J. McLaughlin, J. W. Spatafora & R. Vilgalys. 2006. Reconstructing the early evolution of Fungi using a six-gene phylogeny. Nature 443: 818–822.
[KC01] Kirk, P. M., P. F. Cannon, J. C. David & J. A. Stalpers. 2001. Ainsworth & Bisby’s Dictionary of the Fungi 9th ed. CAB International: Wallingford (UK).
[LK04] Lutzoni, F., F. Kauff, C. J. Cox, D. McLaughlin, G. Celio, B. Dentinger, M. Padamsee, D. Hibbett, T. Y. James, E. Baloch, M. Grube, V. Reeb, V. Hofstetter, C. Schoch, A. E. Arnold, J. Miadlikowska, J. Spatafora, D. Johnson, S. Hambleton, M. Crockett, R. Shoemaker, G.-H. Sung, R. Lücking, T. Lumbsch, K. O’Donnell, M. Binder, P. Diederich, D. Ertz, C. Gueidan, K. Hansen, R. C. Harris, K. Hosaka, Y.-W. Lim, B. Matheny, H. Nishida, D. Pfister, J. Rogers, A. Rossman, I. Schmitt, H. Sipman, J. Stone, J. Sugiyama, R. Yahr & R. Vilgalys. 2004. Assembling the fungal tree of life: progress, classification, and evolution of subcellular traits. American Journal of Botany 91 (10): 1446–1480.