Belongs within: Assamiidae.
The Hypoxestinae are a subfamily of assamiid harvestmen found in Africa and Asia (Santos & Prieto 2010). They are generally characterised by the absence of characters found in other subfamilies and should be considered of questionable monophyly.
Characters (from Santos & Prieto 2010): Anterior margin of carapace unarmed; spiracles visible, unprotected; femur of pedipalp with short ventral teeth; distitarsus I two-segmented; metatarsus II undivided; tarsi III and IV with smooth claws.
<==Hypoxestinae |--Scabrobunus Roewer 1912R35 | `--S. filipes Roewer 1912R35 |--Passula Roewer 1927R35 | `--P. scabricula Roewer 1927R35 |--Podaucheniellus Roewer 1927SP10 | `--*P. bipalaris Roewer 1927S92 |--Nkogoa Roewer 1927SP10 | `--*N. feai Roewer 1927S92 [=N. feaeR35] |--Findia Roewer 1927SP10 | `--*F. atrolutea Roewer 1915S92 |--Congonella Roewer 1935R35 | `--*C. frontalis Roewer 1935R35 |--Randilea Roewer 1935R35 | `--*R. scabricula Roewer 1935R35 [=Hypoxestus scabriculaS92] |--Talaspus Roewer 1935R35 | `--*T. spinimanus Roewer 1935R35 |--Parasesostris Roewer 1915R35 | `--*P. granulatus Roewer 1915S92 |--Metasesostris Roewer 1915R35 | `--*M. armatus Roewer 1915S92 |--Tusipulla Roewer 1935R35 | `--*T. coxalis Roewer 1935R35 |--Podauchenius Sørensen 1896SP10 | `--*P. longipes Sørensen 1896SP10 |--Musola Roewer 1927SP10 | `--*M. longipes Roewer 1927S92 |--Adamauna Roewer 1935SP10 | `--*A. maculatipes Roewer 1935R35 |--Rhabdopygata Roewer 1954SP10, S92 | `--*R. mossambica Roewer 1954S92 |--Lossida Roewer 1935L62, R35 | `--*L. rugosa Roewer 1935R35 |--Lossidacola Roewer 1935L62, R35 | `--*L. pachytarsus Roewer 1935R35 |--Bwitonatus Roewer 1950L62, S92 | `--*B. marlieri Roewer 1950S92 |--Leleupiolus Roewer 1951L62, S92 | `--*L. marmoratus Roewer 1951S92 |--Mecutina Roewer 1935R35 | |--*M. filipes Roewer 1935R35 | `--M. moshinia Roewer 1940S92 |--Sesostranus Roewer 1935R35 | |--*S. niger Roewer 1935R35 | `--S. longipes Roewer 1950S92 |--Bandona Roewer 1927R35 | |--B. boninensis Suzuki 1974S78 | `--B. palpalis Roewer 1927R35 |--Hypoxestinus Roewer 1927SP10 | |--*H. nkogoi Roewer 1927S92 | `--H. frontalis Roewer 1935R35 |--Rhabdopygella Staręga 1992R35, KM20 | |--*R. laevis Roewer 1935S92 | `--R. ferruginea Roewer 1935R35 |--Dongila Roewer 1927SP10 | |--*D. spinosa Roewer 1927S92 | `--D. silvatica Roewer 1927R35 |--Mabwella Roewer 1952L62, S92 | |--*M. wittei Roewer 1952S92 | `--M. trochanteralis Roewer 1954S92 |--Sesostrellus Roewer 1935K85 | |--*S. robustus Roewer 1935R35 | |--S. gibbosus Kauri 1985S92 | `--S. umbonatus (Roewer 1935) [=Sesostris umbonatus]S92 |--Doloressus Roewer 1935 [incl. Ealabius Roewer 1950]K85 | |--*D. cippatus Roewer 1935R35 | |--D. filipes Lawrence 1949S92 | |--D. ghesquierei (Roewer 1950) [=Ealabius ghesquierei]S92 | `--D. palmgreni Kauri 1985S92 |--Dicoryphus Loman 1902R35 [incl. Coelobunus Loman 1902S92, CaelobunusR23, Metarhabdopygus Roewer 1913R35] | |--*D. furvus Loman 1902S92 | |--D. ater (Lawrence 1962) [=Caelobunus ater]S92 | |--D. jeanneli (Roewer 1913) [=Metarhabdopygus jeanneli, Caelobunus jeanneli]S92 | `--D. melanacanthus (Loman 1902) (see below for synonymy)S92 |--Sesostris Sørensen 1910K85 | |--*S. gracilis Sørensen 1910S92 | |--S. brevipes Kauri 1985S92 | |--S. insulanus Roewer 1912K85 | |--S. maculatus Roewer 1915R35 | `--S. maculifer Kauri 1985 [=S. maculiferus]S92 |--Viglua Roewer 1940SP10 | |--*V. brunnipes Roewer 1940S92 | `--V. machadoi Lawrence 1949S92 | |--V. m. machadoiS92 | |--V. m. granulosa Lawrence 1957S92 | `--V. m. major Lawrence 1949 [=V. majora]S92 |--Spinixestus Roewer 1952K85 | |--*S. armatus Roewer 1952S92 | |--S. benderanus Kauri 1985S92 | |--S. leleupi Kauri 1985S92 | |--S. polycuspidatus Kauri 1985S92 | |--S. rufus Roewer 1952S92 | `--S. siteteus Roewer 1961S92 |--Rhabdopygus Roewer 1912SP10 | |--*R. fuscus Roewer 1912SP10 | |--R. benoiti Kauri 1985SP10 | |--R. funilignorum Santos & Prieto 2010SP10 | |--R. maculatus Roewer 1935SP10 | |--R. puylaerti Bauer & Prieto 2009SP10 | |--R. robustus Lawrence 1957SP10 | |--R. rugipalpis Roewer 1952SP10 | `--R. termitarum Roewer 1951SP10 `--Hypoxestus Loman 1902S92 [=Amasilla Strand 1911R23, Amasis Sørensen 1910 non Leach 1817R23] | i. s.: H. bituberculatus Lawrence 1962S92 | H. coxicornis Roewer 1940S92 | H. patellaris (Sørensen 1910)S92 [=*Amasis patellarisKM20] | H. planus Goodnight & Goodnight 1959S92 | H. roeweri Staręga 1992 [=Randilea obscura Roewer 1952 non H. obscurus Roewer 1912]S92 | H. trituberculatus Lawrence 1962S92 |--*H. (Hypoxestus) levis Loman 1902 (see below for synonymy)S92 `--H. (Lomanus Kauri 1985)K85 |--H. (*L.) glaber Kauri 1985K85 |--H. (L.) ealanus Kauri 1985K85 `--H. (L.) scaphoides Kauri 1985K85
Dicoryphus melanacanthus (Loman 1902) [=Coelobunus melanacanthus, Caelobunus melanacanthus; incl. Co. fuscus Roewer 1912, Ca. fuscus]S92
*Hypoxestus (Hypoxestus) levis Loman 1902 [incl. Amasis accentuatus Sørensen 1910, H. accentuatus, A. mesoleucus Sørensen 1910, H. mesoleucus, H. obscurus Roewer 1912, Amasis quadricornis Sørensen 1910, H. quadricornis]S92
*Type species of generic name indicated
References
[K85] Kauri, H. 1985. Opiliones from central Africa. Koninklijk Museum voor Midden-Afrika, Turveren, België, Zoologische Wetenschappen, Annalen 245: 1–167.
[KM20] Kury, A. B., A. C. Mendes, L. Cardoso, M. S. Kury & A. A. Granado. 2020. WCO-Lite: Online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0—Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Published by the author: Rio de Janeiro.
[L62] Lawrence, R. F. 1962. Resultats scientifiques des missions zoologiques de l’I.R.S.A.C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXIV. Opiliones. Annales du Musée Royal de l’Afrique Centrale: Sciences Zoologiques 110: 9–89.
[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.
[R35] Roewer, C.-F. 1935. Alte und neue Assamiidae. Weitere Weberknechte VIII. (8. Ergänzung der “Weberknechte der Erde” 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Uebersee-Museum in Bremen 1: 1–168.
[SP10] Santos, R., & C. E. Prieto. 2010. Los Assamiidae (Opiliones: Assamiidae) de Río Muni (Guinea Ecuatorial), con la descripción de ocho nuevas especies. Revista de Biologia Tropical 58 (1): 203–243.
[S92] Staręga, W. 1992. An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum 33 (2): 271–336.
[S78] Suzuki, S. 1978. Three harvestmen (Arachnida, Opiliones) from the Bonin Islands. Annotationes Zoologicae Japonenses 51 (3): 179–185.