Iocheirata

Geogarypus macrodactylus, from Harms & Dunlop (2017).

Belongs within: Pseudoscorpiones.
Contains: Panctenata, Neobisiidae.

The Iocheirata are a major clade of pseudoscorpions characterised by the presence of venom glands in the pedipalpal chelae (Murienne et al. 2008).

Synapomorphies (Murienne et al. 2008): Venom glands present in pedipalpal chela; posterior maxillary lyrifissure present; trichobothrium isb on external face of fixed chelal finger; slit sensillum of legs I and II subdistally placed.

<==Iocheirata [Diplosphyronida, Garypoidea, Hemictenata, Mestommatina, Neobisiinea, Neobisioidea, Syarinidae]
    |  i. s.: Hyarinus hesperusM90
    |         Aglaochitra rexM90
    |         Microblothrus Mahnert 1985CH21a
    |--+--IdeoroncidaeMHG08
    |  |    |--Pseudalbiorix veracruzensisMHG08
    |  |    |--AlbiorixM90
    |  |    |--IdeoroncusM01
    |  |    |    |--I. anophthalmusM01
    |  |    |    |--I. cavicola Mahnert 2001M01
    |  |    |    |--I. divisusM01
    |  |    |    `--I. setosus Mahnert 1984M01
    |  |    |--DhanusJ97
    |  |    |    |--D. doveri Bristowe 1952J97
    |  |    |    `--D. siamensis (With 1906) [=Ideoroncus siamensis]J97
    |  |    `--Negroroncus longedigitatus Beier 1944J97
    |  `--Bochicidae [Vachoniidae]MHG08
    |       |--LeucohyinaeMHG08
    |       `--BochicinaeMHG08
    |            |--Bochica withi (Chamberlin 1923) [=Ideoroncus withi]J97
    |            |--AntillobisiumM01
    |            |--TroglobochicaM01
    |            `--Spelaeobochica Mahnert 2001M01
    |                 `--*S. allodentatus Mahnert 2001M01
    `--+--Indohya [Hyidae]MHG08
       |    |--I. damocles Harvey & Volschenk 2007EH08
       |    |--I. gollum Harvey & Volschenk 2007EH08
       |    |--I. humphreysi (Harvey 1993)EH08
       |    `--I. napierensis Harvey & Volschenk 2007EH08
       `--+--PanctenataMHG08
          `--+--IdeobisiinaeMHG08
             |    |--+--Nannobisium beieri Mahnert 1979MHG08, M01
             |    |  `--GeogarypidaeJ97
             |    |       |--Geogarypus Chamberlin 1930MHG08, CH21b [incl. Indogarypus Beier 1957CH21b]
             |    |       |    |--*G. minor (Koch 1873) [=Garypus minor]CH21b
             |    |       |    |--G. connatus Harvey 1986CH21b
             |    |       |    |--G. elegans (With 1906) [=Garypus elegans, Ga. personatus Simon 1901 non Simon 1900]J97
             |    |       |    |--G. exochus Harvey 1986CH21b
             |    |       |    |--G. facetus Cullen & Harvey 2021CH21b
             |    |       |    |--G. indicus (Beier 1930) [=Garypus indicus, Ge. (*Indogarypus) indicus]CH21b
             |    |       |    |--G. javanusH83
             |    |       |    |--G. longidigitatus (Rainbow 1897)CH21b
             |    |       |    |--G. macrodactylus Beier 1937S93
             |    |       |    |--G. maculatus With 1907J97
             |    |       |    |--G. major Beier 1937S93
             |    |       |    |--G. personatus (Simon 1900) [=Garypus personatus]J97
             |    |       |    |--G. pisinnus Harvey 1986CH21b
             |    |       |    |--G. plusculus Cullen & Harvey 2021CH21b
             |    |       |    |--G. rhantus Harvey 1981CH21b
             |    |       |    |--G. taylori Harvey 1986CH21b
             |    |       |    `--G. tenuis Chamberlin 1930J97
             |    |       `--Afrogarypus Beier 1931CH21b
             |    |            `--A. seychellesensis (Beier 1940) [=Geogarypus seychellesensis]J97
             |    `--+--Ideobisium Balzan 1891MHG08, S06 [incl. Xenobisium Muchmore 1982 (n. n.)J97]
             |       |    |--I. antipodum (Simon 1880) [=Obisium antipodum]H81
             |       |    |--I. balzanii With 1905J97
             |       |    |--I. bipectinatumH83
             |       |    |--I. peregrinum Chamberlin 1930J97
             |       |    `--I. theveneti (Simon 1878)S06
             |       `--+--Parahya Beier 1957MHG08, H09 [Parahyidae]
             |          |    `--P. submersa (Bristowe 1931) [=Obisium submersum]J97
             |          `--Ideoblothrus Balzan 1892MHG08, HL08a [incl. Pachychitra Chamberlin 1938HL08a]
             |               |--*I. similis (Balzan 1892) [=Ideobisium (*Ideoblothrus) similis]HL08a
             |               |--I. descartes Harvey & Edward 2007HL08a
             |               |--I. floridensisM90
             |               |--I. linnaei Harvey & Leng 2008HL08a
             |               |--I. maya (Chamberlin 1938) [=*Pachychitra maya]HL08a
             |               |--I. milikapiti Harvey & Edward 2007HL08a
             |               |--I. nesotymbus Harvey & Edward 2007T13
             |               |--I. papillon Harvey 1991HL08a
             |               |--I. pisolitus Harvey & Edward 2007HL08a
             |               |--I. pugil (Beier 1964) [=Ideobisium pugil]J97
             |               |    |--I. p. pugilJ97
             |               |    `--I. p. robustum (Beier 1966) [=Ideobisium pugil robustum]J97
             |               |--I. seychellesensis (Chamberlin 1930) [=Ideobisium seychellesensis]J97
             |               |--I. westi Harvey & Edward 2007HL08a
             |               `--I. woodi Harvey 1991H91
             `--+--+--Mirobisium [Gymnobisiidae]MHG08
                |  `--Chitrella [Chitrellinae]MHG08
                |       |--C. calaMHG08
                |       `--C. cavicolaM90
                `--NeobisiidaeMHG08

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