
Belongs within: Pseudoscorpiones.
Contains: Panctenata, Neobisiidae.
The Iocheirata are a major clade of pseudoscorpions characterised by the presence of venom glands in the pedipalpal chelae (Murienne et al. 2008).
Synapomorphies (Murienne et al. 2008): Venom glands present in pedipalpal chela; posterior maxillary lyrifissure present; trichobothrium isb on external face of fixed chelal finger; slit sensillum of legs I and II subdistally placed.
<==Iocheirata [Diplosphyronida, Garypoidea, Hemictenata, Mestommatina, Neobisiinea, Neobisioidea, Syarinidae]
| i. s.: Hyarinus hesperusM90
| Aglaochitra rexM90
| Microblothrus Mahnert 1985CH21a
|--+--IdeoroncidaeMHG08
| | |--Pseudalbiorix veracruzensisMHG08
| | |--AlbiorixM90
| | |--IdeoroncusM01
| | | |--I. anophthalmusM01
| | | |--I. cavicola Mahnert 2001M01
| | | |--I. divisusM01
| | | `--I. setosus Mahnert 1984M01
| | |--DhanusJ97
| | | |--D. doveri Bristowe 1952J97
| | | `--D. siamensis (With 1906) [=Ideoroncus siamensis]J97
| | `--Negroroncus longedigitatus Beier 1944J97
| `--Bochicidae [Vachoniidae]MHG08
| |--LeucohyinaeMHG08
| `--BochicinaeMHG08
| |--Bochica withi (Chamberlin 1923) [=Ideoroncus withi]J97
| |--AntillobisiumM01
| |--TroglobochicaM01
| `--Spelaeobochica Mahnert 2001M01
| `--*S. allodentatus Mahnert 2001M01
`--+--Indohya [Hyidae]MHG08
| |--I. damocles Harvey & Volschenk 2007EH08
| |--I. gollum Harvey & Volschenk 2007EH08
| |--I. humphreysi (Harvey 1993)EH08
| `--I. napierensis Harvey & Volschenk 2007EH08
`--+--PanctenataMHG08
`--+--IdeobisiinaeMHG08
| |--+--Nannobisium beieri Mahnert 1979MHG08, M01
| | `--GeogarypidaeJ97
| | |--Geogarypus Chamberlin 1930MHG08, CH21b [incl. Indogarypus Beier 1957CH21b]
| | | |--*G. minor (Koch 1873) [=Garypus minor]CH21b
| | | |--G. connatus Harvey 1986CH21b
| | | |--G. elegans (With 1906) [=Garypus elegans, Ga. personatus Simon 1901 non Simon 1900]J97
| | | |--G. exochus Harvey 1986CH21b
| | | |--G. facetus Cullen & Harvey 2021CH21b
| | | |--G. indicus (Beier 1930) [=Garypus indicus, Ge. (*Indogarypus) indicus]CH21b
| | | |--G. javanusH83
| | | |--G. longidigitatus (Rainbow 1897)CH21b
| | | |--G. macrodactylus Beier 1937S93
| | | |--G. maculatus With 1907J97
| | | |--G. major Beier 1937S93
| | | |--G. personatus (Simon 1900) [=Garypus personatus]J97
| | | |--G. pisinnus Harvey 1986CH21b
| | | |--G. plusculus Cullen & Harvey 2021CH21b
| | | |--G. rhantus Harvey 1981CH21b
| | | |--G. taylori Harvey 1986CH21b
| | | `--G. tenuis Chamberlin 1930J97
| | `--Afrogarypus Beier 1931CH21b
| | `--A. seychellesensis (Beier 1940) [=Geogarypus seychellesensis]J97
| `--+--Ideobisium Balzan 1891MHG08, S06 [incl. Xenobisium Muchmore 1982 (n. n.)J97]
| | |--I. antipodum (Simon 1880) [=Obisium antipodum]H81
| | |--I. balzanii With 1905J97
| | |--I. bipectinatumH83
| | |--I. peregrinum Chamberlin 1930J97
| | `--I. theveneti (Simon 1878)S06
| `--+--Parahya Beier 1957MHG08, H09 [Parahyidae]
| | `--P. submersa (Bristowe 1931) [=Obisium submersum]J97
| `--Ideoblothrus Balzan 1892MHG08, HL08a [incl. Pachychitra Chamberlin 1938HL08a]
| |--*I. similis (Balzan 1892) [=Ideobisium (*Ideoblothrus) similis]HL08a
| |--I. descartes Harvey & Edward 2007HL08a
| |--I. floridensisM90
| |--I. linnaei Harvey & Leng 2008HL08a
| |--I. maya (Chamberlin 1938) [=*Pachychitra maya]HL08a
| |--I. milikapiti Harvey & Edward 2007HL08a
| |--I. nesotymbus Harvey & Edward 2007T13
| |--I. papillon Harvey 1991HL08a
| |--I. pisolitus Harvey & Edward 2007HL08a
| |--I. pugil (Beier 1964) [=Ideobisium pugil]J97
| | |--I. p. pugilJ97
| | `--I. p. robustum (Beier 1966) [=Ideobisium pugil robustum]J97
| |--I. seychellesensis (Chamberlin 1930) [=Ideobisium seychellesensis]J97
| |--I. westi Harvey & Edward 2007HL08a
| `--I. woodi Harvey 1991H91
`--+--+--Mirobisium [Gymnobisiidae]MHG08
| `--Chitrella [Chitrellinae]MHG08
| |--C. calaMHG08
| `--C. cavicolaM90
`--NeobisiidaeMHG08