Laphriinae

Belongs within: Asilidae.
Contains: Atomosini.

The Laphriinae are a group of robber flies lacking a conspicuous stylus on the antenna (Wood 1981). Members of the genus Laphria are particularly large, hairy robber flies that mimic bees in appearance. The Andrenosomatini are characterised by a dorsoventrally strongly flattened proboscis and flat, lamellate palpi, and females bear a cylindrical anal tube formed from the cerci (Fisher 2009).

Characters (from Wood 1981): Antenna blunt apically, without stylus, or with one or two abruptly tapered short microsegments. Palpus two-segmented. Wing with R₂₊₃ joining R₁ proximal to end of R₁, with cell r thus separated from wing margin; crossvein m-cu present; thus apex of cell bm closed by three veins. Male terminalia rotated 180° permanently.

<==LaphriinaeF09
    |  i. s.: Smeryngolaphria [Dasytrichini]F09
    |           `--S. numitorF09
    |--AndrenosomatiniF09
    |    |--CerotainiopsF09
    |    |--ExodasyllisF09
    |    |--Neophoneus [Neophoneini]F09
    |    |    `--N. amandusF09
    |    |--PilicaF09
    |    |    |--*P. formidolosaF09
    |    |    `--P. erythrogasterF09
    |    |--DasandraF09
    |    |    |--*D. ayalaiF09
    |    |    |--D. hespenheideiF09
    |    |    `--D. zanutoiF09
    |    |--DasyllisF09
    |    |    |--D. coralliaF09
    |    |    |--D. funebrisF09
    |    |    |--D. haemorrhoaF09
    |    |    `--D. phoenicogasterF09
    |    |--EurycheilisF09
    |    |    |--E. bromleyiF09
    |    |    |--E. colliniF09
    |    |    |--E. eufriesiaeF09
    |    |    `--E. nigraF09
    |    `--AndrenosomaBT04
    |         |--*A. (Andrenosoma) atrumF09
    |         |--A. cinereumF09
    |         |--A. fulvicaudumF09
    |         |--A. (Pogonosoma) ridingsiW81
    |         |--A. sarcophagumF09
    |         `--A. xanthocnemumF09
    `--+--AtomosiniBT04
       `--Laphriini [Dasylechiini]BT04
            |--MairaBT04
            |--DasylechiaW81
            |--Ichneumolaphria fascipennisF09
            |--*Phellopteron farriF09
            |--OrthogonisW81
            |    |--O. christmasensis (Daniels 2012) (see below for synonymy)JG19
            |    `--O. stygiaW81
            |--RhopalogasterF09
            |    |--R. araujoiF09
            |    `--R. micronyxF09
            |--LampriaF09
            |    |--L. aurifexF09
            |    |--L. clavipesF09
            |    `--L. rubriventrisW81
            `--LaphriaBT04
                 |--L. atraR26
                 |--L. aureaR26
                 |--L. bomboides Macquart 1847E12
                 |--L. coerulea Meigen in Boisduval 1835B35
                 |--L. colorata Boisduval 1835B35
                 |--L. flavaG20
                 |--L. gilvaK01
                 |--L. hirtaC91
                 |--L. maroccanaL49
                 |--L. nigrocaerulea (van der Wulp 1872)JG19
                 |--L. sadalesW81
                 `--L. thoracicaCW09

Orthogonis christmasensis (Daniels 2012) [=Laphria christmasensis, L. nigrocaerulea Kirby 1889 non (van der Wulp 1872)]^JG19

*Type species of generic name indicated

References

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[BT04] Bybee, S. M., S. D. Taylor, C. R. Nelson & M. F. Whiting. 2004. A phylogeny of robber flies (Diptera: Asilidae) at the subfamilial level: molecular evidence. Molecular Phylogenetics and Evolution 30: 789–797.

[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 1. Melbourne University Press: Carlton (Victoria).

[CW09] Cumming, J. M., & D. M. Wood. 2009. Adult morphology and terminology. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 9–50. NRC Research Press: Ottawa.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F09] Fisher, E. M. 2009. Asilidae (robber flies, assassin flies, moscas cazadoras, moscas ladronas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 585–632. NRC Research Press: Ottawa.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[K01] Kertész, K. 1901. Legyek [Dipteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 179–201. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[W81] Wood, G. C. 1981. Asilidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 549–573. Research Branch, Agriculture Canada.