Belongs within: Lauxanioidea.
Contains: Trigonometopini.
The Lauxaniidae are a family of flies whose larvae mostly live in fallen leaves and other rotting vegetation. Several genera include nidicolous species, feeding on decaying matter in bird nests. Adults graze on fungi growing on fallen leaves (Gaimari & Silva 2010).
The internal classification of the family is somewhat poorly established. The Eurychoromyiinae have previously been recognised as a separate family due to a reduction in setation but examination at high magnification has found that a number of ‘lost’ setae are present but greatly reduced. The Homoneurinae are characterised by wings with costal setulae reaching the apex of vein R4+5 whereas in Lauxaniinae they end shortly past vein R2+3; however, it seems likely that the ‘Lauxaniinae’ are paraphyletic to the Homoneurinae. Similarly, certain near-cosmopolitan genera such as Sapromyza and Homoneura may prove to comprise multiple distinct lineages when further revised (Gaimari & Silva 2010).
Characters (from Colless & McAlpine 1991): One or two fronto-orbital bristles; costa unbroken; Sc complete; Cu+1A discontinued well before margine; vein 2A generally visible beyond alula; anepisternite with one or two mesopleural bristles; tibiae with preapical dorsal bristles; sternite 1 well developed, sclerotised.
<==Lauxaniidae [Sapromyzidae] |--LauxaniinaeGS10 | | i. s.: LauxaniaGS10 | | |--L. aeneaRD77 | | |--L. cylindricornis [=Micropeza (Lauxania) cylindricornis]G20 | | `--L. rufitarsisR26 | `--TrigonometopiniP07 |--SapromyzaGS10 [SapromyzinaeK01] | |--S. flavimanaCM91 | |--S. frontalisGS10 | |--S. lupulinaK01 | |--S. nigransGS10 | |--S. picticornisGS10 | |--S. rotundicornisS87 | |--S. slossonaeGS10 | |--S. sororiaGS10 | `--S. vinnulaGS10 |--HomoneurinaeGS10 | |--TrypetisomaGS10 | | |--T. sticticumGS10 | | `--T. zacatecasenseGS10 | `--HomoneuraGS10 | | i. s.: H. bispinaGS10 | | H. proximellaCM91 | |--H. (Homoneura) philadelphicaGS10 | `--H. (Tarsohomoneura)GS10 | |--H. (T.) americanaGS10 | `--H. (T.) ornatipesGS10 `--Eurychoromyiinae [Eurychoromyiidae]GS10 |--EurychoromyiaGS10 |--Tauridion shewelliGS10 |--Choryeuromyia senismaGS10 |--EurystratiomyiaGS10 |--RoryeuchomyiaGS10 |--EuryhendelimyiaGS10 `--PhysegeniopsisGS10 |--P. albetoGS10 |--P. ankhoideaGS10 `--P. hadrocaraGS10 Lauxaniidae incertae sedis: NoonamyiaO98 [NoonamyiiniP07] Paranomina Hendel 1907P07 LyciellaS87 |--L. decipiensWT11 `--L. discolorS87 CerastocaraZS10 CephaloconusZS10 PoecilohetarusZS10 NeogriphoneuraGS10 |--N. laevifronsGS10 |--N. sordidaGS10 |--N. striatifronsGS10 |--N. strigaGS10 |--N. tertiaGS10 `--N. timidaGS10 Siphonophysa albipesGS10 NeodeceiaGS10 |--N. cineraceaGS10 `--N. exulGS10 HypagogaGS10 Oncodometopus umbrosusGS10 TricholauxaniaGS10 PseudocalliopeGS10 |--P. eucephalaGS10 |--P. flavicepsGS10 `--P. longicornisS87 Poecilominettia maculaGS10 MeiosimyzaGS10 Calliopum quadrisetosumGS10 PseudogriphoneuraGS10 Camptoprosopella borealisGS10 Poecilolycia serrataGS10 MelanomyzaGS10 |--M. (Melanomyza)GS10 | |--M. (M.) gracilipesGS10 | `--M. (M.) proanaGS10 `--M. (Lauxaniella)GS10 |--‘Lauxaniella’ femoralisM81 `--M. (L.) manuleataGS10 TrivialiaGS10 |--T. celesteGS10 |--T. fuscocapitataGS10 |--T. nigroapicataGS10 `--T. puellaGS10 XenochaetinaGS10 |--X. flavipennisGS10 |--X. muscariaGS10 `--X. pallidaGS10 PhysoclypeusGS10 |--P. coquillettiGS10 |--P. farinosusGS10 |--P. hendeliGS10 |--P. unimaculatusGS10 `--P. zebrinusGS10 Xeniconeura costalisGS10 Trisapromyza vittigeraGS10 AllominettiaGS10 |--A. approximataGS10 |--A. assimilisGS10 `--A. genisetaGS10 Mallochomyza citreifronsGS10 Steganolauxania latipennisGS10 Cephalella fumipennisGS10 TriconopsisGS10 AsilostomaGS10 |--A. atricepsGS10 |--A. flavifaciesGS10 |--A. pallipesGS10 `--A. palpalisGS10 PachyopellaGS10 |--P. flavidaGS10 `--P. ornataGS10 StenolauxaniaGS10 |--S. nubeculaGS10 `--S. striataGS10 Minettia [Minettiinae]GS10 |--M. aibonitoGS10 |--M. flaveolaGS10 |--M. hubbardiiGS10 `--M. lupulinaGS10 XenopterellaGS10 |--X. beameriGS10 `--X. obliquaGS10 BaliopteridionGS10 HirtodeceiaGS10 ParacestrotusGS10 DeutominettiaGS10 |--D. bimaculataGS10 `--D. genisetaS87 ScutominettiaGS10 |--S. innubaGS10 `--S. mallochiGS10 GriphoneuraGS10 ChaetominettiaGS10 |--C. corollaeGS10 `--C. spinitibiaGS10 DryomyzotheaGS10 SetulinaGS10 |--S. geminataGS10 |--S. interjectaGS10 |--S. primaGS10 `--S. stataGS10 ChaetocoeliaGS10 |--C. calopteraGS10 |--C. distinctissimaGS10 `--C. pallansGS10 DryosapromyzaGS10 SciosapromyzaGS10 |--S. coronataGS10 |--S. ficulneaGS10 `--S. tinctinervisGS10 Pseudominettia platypezaGS10 AgriphoneuraGS10 EriurgusGS10 OcellominettiaGS10 CelypholauxaniaGS10 Procrita pectinataGS10 BlepharolauxaniaGS10 Platygraphium penicillatumGS10 Freyia nigritaGS10 PhysegenuaGS10 Neoxangelina ornataGS10 Minilauxania bulbifaciesGS10 RitaemyiaGS10 ScutolauxaniaGS10 NeopachycerinaGS10 GibbolauxaniaGS10 |--G. alienaGS10 `--G. lateralisGS10 DeceiaGS10 Marmarodeceia marmorataGS10 Neominettia contiguaGS10 Depressa striatipennisCM91 ‘Lonchaea’ senescensS87 Eolausanites Hong 2002H02 `--*E. ellipticus (Hong 1981) [=Trypaneoides ellipticus]H02 ChamaelauxaniaP92 HemilauxaniaP92
*Type species of generic name indicated
References
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[GS10] Gaimari, S. D., & V. C. Silva. 2010. Lauxaniidae (lauxaniid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 971–995. NRC Research Press: Ottawa.
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[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.
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[ZS10] Zborowski, P., & R. Storey. 2010. A Field Guide to Insects in Australia 3rd ed. Reed New Holland: Sydney.