Loeblichiidae

Axial (left) and equatorial sections of Lysella gadukensis, from Cózar & Vachard (2001).

Belongs within: Endothyroida.
Contains: Dainella, Bessiella, Paradainella, Endothyranopsidae, Loeblichiinae, Fusulinoida, Paralysella, Eostaffellidae, Reichelinidae.

The Loeblichiidae are a lineage of fusulinid Foraminifera from the lower Carboniferous with numerous chambers that were regarded by Cózar & Vachard (2001) as including the ancestors of later fusulinids from the later Carboniferous and Permian. The basal ‘Spinoendothyrinae’ possessed numerous rather inflated chambers containing secondary deposits in the form of crustae or spines but lacking chomata. True chomata were evolved by the more derived Dainellinae (Cózar & Vachard 2001).

The Ozawainellidae are a group of fusulinids known from the upper Lower Carboniferous to the Upper Permian (Loeblich & Tappan 1964).

<==Loeblichiidae [Fusulinacea, Loeblichinidae, Spinoendothyrinae]
|--Inflatoendothyra Brazhnikova & Vdovenko in Vdovenko 1972CV01
| `--*I. inflata (Lipina 1955) (see below for synonymy)R-CB96
|--Elergella Conil in Shilo et al. 1984CV01
| `--*E. simakovi Conil 1984R-CB96
`--+--Spinoendothyra Lipina in Anonymous 1963CV01
| `--*S. costifera (Lipina 1955) [=Endothyra costifera]R-CB96
|--+--Spinobrunsiina Conil & Longerstaey in Conil et al. 1980CV01
| `--Pseudoplanoendothyra Brazhnikova & Vdovenko in Brazhnikova 1982CV01
| `--*P. rotayi (Brazhnikova 1962) [=Quasiendothyra rotayi]R-CB96
|--+--Banffella Mamet in Mamet & Skipp 1970CV01
| `--Urbanella Malakhova in Anonymous 1963 [Urbanellinae]CV01
| `--*U. urbana (Malakhova 1954) [=Quasiendothyra urbana]R-CB96
`--+--Pseudochernyshinella Brazhnikova 1974CV01
| `--*P. biformis [=Endothyra subrotunda biformis]R-CB96
`--Dainellinae [Dainellidae]CV01
|--DainellaCV01
|--+--BessiellaCV01
| `--Vissarionovella Cózar & Vachard 2001CV01
| |--*V. tujmasensis (Vissarionova 1948) (see below for synonymy)CV01
| |--V. donzellii Cózar & Vachard 2001CV01
| `--V. llangollensis (Conil & Ramsbottom in Conil et al. 1981) [=Bessiella llangollensis]CV01
|--+--ParadainellaCV01
| |--+--EndothyranopsidaeCV01
| | `--GloboendothyridaeM13
| | |--EblanainaeM13
| | `--GloboendothyrinaeM13
| | |--Carbotarima Brenckle 2004M13
| | |--Planogloboendothyra Hance, Hou & Vachard 2011M13
| | `--Globoendothyra Reitlinger 1959R-CB96
| | | i. s.: G. baileyiCM01
| | | G. tomiliensisR79
| | |--*G. (Globoendothyra) globulus (Moeller 1878)R-CB96 (see below for synonymy)
| | `--G. (Eogloboendothyra Vdovenko 1972)R-CB96, CV01, R-CB96
| | `--G. (*E.) ‘parva’ (Tchernysheva 1948) (see below for synonymy)R-CB96
| `--+--LoeblichiinaeCV01
| |--+--FusulinoidaCV01
| | `--Eoparastaffellina Vdovenko 1971CV01, R-CB96
| | |--*E. subglobosa (Vdovenko 1971) [=Eoparastaffella (*Eoparastaffellina) subglobosa]R-CB96
| | `--E. umbilicata (Bozorgnia 1973) [=Lysella umbilicata]CV01
| `--Eoparastaffella Vdovenko 1954LT64 [incl. Bozorgniella Cózar & Vachard 2001CV01]
| |--*E. simplex (Vdovenko 1954)R-CB96 (see below for synonymy)
| |--‘Dainella’ tumultuosa Bozorgnia 1973 [=*Bozorgniella tumultuosa]CV01
| |--‘Pseudoendothyra’ florigena Pronina 1963 [=Bozorgniella florigena, Eoparastaffella florigena]CV01
| |--‘Eoparastaffella’ fundata Simonova 1975 [=Bozorgniella fundata]CV01
| |--‘Plectogyra’ librans Conil & Lys 1964 [=Bozorgniella librans]CV01
| |--‘Eoparastaffella (Eoparastaffellina)’ rotunda Vdovenko 1971 [=Bozorgniella rotunda]CV01
| `--‘Eoparastaffella’ tumida Pronina 1963 [=Bozorgniella tumida, Pseudoendothyra tumida]CV01
`--+--Lysella Bozorgnia 1973CV01
| |--*L. gadukensis Bozorgnia 1973 [incl. L. conferta Bozorgnia 1973]CV01
| |--L. amenta (Ganelina 1966) [=Dainella amenta]CV01
| |--L. compacta (Postoyalko 1970) [=Dainella compacta]CV01
| |--L. incomposita (Postoyalko 1970) [=Dainella incomposita]CV01
| |--L. magna (Vdovenko 1954) [=Quasiendothyra magna]CV01
| |--L. manifesta (Ganelina 1966) [=Dainella manifesta]CV01
| |--L. mediocriformisCV01
| |--L. micula (Postoyalko 1970) [=Dainella micula]CV01
| `--L. uralica (Postoyalko 1970) [=Dainella uralica]CV01
`--+--ParalysellaCV01
`--Ozawainellida [Ozawainelloida]M13
| i. s.: Hubeiella Lin 1977R-CB96
| `--*H. simplex Lin 1977R-CB96
|--EostaffellidaeR-CB96
|--ReichelinidaeM13
`--Ozawainellidae [Ozawainellacea, Ozawainellinae]CV01
|--Ozawainella Thompson 1935LT64
| `--*O. angulata (Colani 1924) [=Fusulinella angulata]LT64
|--Eostaffelloides Miklukho-Maclay 1959HW93
| `--*E. orientalis Miklukho-Maclay 1959R-CB96
|--Kangvarella Saurin 1962HW93
| `--*K. irregularis Saurin 1962R-CB96
`--Paramillerella Thompson 1951LT64
|--*P. advena (Thompson 1944) [=Millerella advena]LT64
|--P. amplaLT64
|--P. circuliLT64
`--P. pinguisLT64

*Eoparastaffella simplex (Vdovenko 1954)R-CB96 [=Parastaffella (Eoparastaffella) simplexR-CB96, Bozorgniella simplexCV01]

Globoendothyra (*Eogloboendothyra) ‘parva’ (Tchernysheva 1948) [=Endothyra globulus var. parva non E. parva Möller 1879]R-CB96

*Globoendothyra (Globoendothyra) globulus (Moeller 1878)R-CB96 [=Endothyra globulusR-CB96, *G. pseudoglobulus Reitlinger 1959R-CB96, Endothyranopsis pseudoglobulusLT64]

*Inflatoendothyra inflata (Lipina 1955) [=Spinoendothyra (*Inflatoendothyra) inflata, Endothyra oldalipinae Loeblich & Tappan 1987]R-CB96

*Vissarionovella tujmasensis (Vissarionova 1948) [=Eostaffella tujmasensis, Dainella tujmasensis, Lysella tujmasensis]CV01

*Type species of generic name indicated

References

[CM01] Cózar, P., & B. L. Mamet. 2001. Planohowchinia (Lasiodiscidae, Foraminiferida), a new Late Viséan genus, southwestern Spain. Journal of Foraminiferal Research 31 (3): 228–232.

[CV01] Cózar, P., & D. Vachard. 2001. Dainellinae subfam. nov. (Foraminiferida du Carbomifère inférieur), révision et nouveaux taxons. Geobios 34 (5): 505–526.

[HW93] Hart, M. B., & C. L. Williams. 1993. Protozoa. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 43–70. Chapman & Hall: London.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[M13] Mikhalevich, V. I. 2013. New insight into the systematics and evolution of the Foraminifera. Micropaleontology 59 (6): 493–527.

[R-CB96] Rauzer-Chernousova, D. M., F. R. Bensh, M. V. Vdovenko, N. B. Gibshman, E. Y. Leven, O. A. Lipina, E. A. Reitlinger, M. N. Solovieva & I. O. Chedija. 1996. Spravočnik po Sistematike Foraminifer Paleozoâ (Èndotiroidy, Fuzulinoidy). Rossijskaâ Akademiâ Nauk, Geologičeskij Institut, Moskva “Nauka”.

[R79] Ross, C. A. 1979. Carboniferous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A254–A290. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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