Madarina

Belongs within: Conoderinae.

The Madarina are baridine weevils with uninterrupted surfaces between prosternum, meso- and metaventrite, and an oblique, dorsally visible pygidium (Legalov 2018). They are usually elliptical or rhomboidal in overall shape, and often brightly coloured. Members of the genus Microstrates are found in palm flowers.

Leptoschoinus and related genera have historically been separated by their relatively short body into a subtribe Leptoschoinina. However, Legalov (2018) did not regard this distinction as worthy of such recognition. Hustache (1951) established the genus Leptoschoinella for species from South America differing from Leptoschoinus in having more closely spaced fore coxae and a more elongate body shape.

<==Madarina [Leptoschoinides, Leptoschoinina, Thaliabaridina]L18
    |--Parapiperis minutus Hustache 1951H51
    |--Acanthomadarus amplicollis Hustache 1951H51
    |--Parasolaria ferruginipes Hustache 1951H51
    |--Solariopsis punctaticollis Hustache 1951H51
    |--Acythopeus Pascoe 1874D09
    |--Eumycterus albosquamulatus Boheman 1838D09
    |--Zena virgata Hustache 1938D09
    |--Stictobaris Casey 1892A02
    |    `--S. cribrata (LeConte 1876)D09
    |--Neomadarus Hustache 1938D09
    |    `--N. saba (Fabricius 1801) [incl. Sibariops boliviensis Hustache 1924, S. fuirenae Bondar 1943]K83
    |--Amadarellus Hustache 1951H51
    |    `--*A. bondari Hustache 1951H51
    |--Apolpones Hustache 1951H51
    |    `--*A. rectirostris Hustache 1951H51
    |--Megabaris Hustache 1951H51
    |    `--*M. quadriguttata (Klug 1829)H51, D09 [=Centrinus quadriguttoctusH51]
    |--Thaliabaris Bondar 1943D09, K83
    |    `--T. inermis Marshall 1952D09
    |--Leptoschoinus Dejean 1836L18
    |    `--L. fucatus (Klug 1829)D09
    |--Glyptobaris Casey 1892A02
    |    |--G. lecontei Champion 1909D09
    |    `--G. liturata (Fabricius 1801) [incl. G. amnicola Casey 1922]K83
    |--MicrorhinusD09
    |    |--M. cuneirostris (Casey 1922) [incl. M. ellipticus Casey 1922]K83
    |    `--M. mutilus (Boheman 1844)D09
    |--AthesapeutaD09
    |    |--A. cyperiB88
    |    `--A. vinculata Faust 1894D09
    |--Onychobaris LeConte 1876A02
    |    |--O. alutacea Hustache 1951H51
    |    `--O. densa LeConte 1876D09
    |--BrachybarisH51
    |    |--‘Baridius’ axillarisH51
    |    `--B. grandis Hustache 1951H51
    |--LoboderesD09
    |    |--L. citriventris Hustache 1938D09
    |    `--L. longicollis Hustache 1951H51
    |--Epipedomorphus Hustache 1951L18, H51
    |    |--*E. argentinensis Hustache 1951H51
    |    `--E. guyanensis Hustache 1951H51
    |--TripusidiaH51
    |    |--T. punctulataH51
    |    |--T. sparsepunctata Hustache 1951H51
    |    `--T. subglabra Hustache 1951H51
    |--SolariaD09
    |    |--S. amazonica [=Onychobaris amazonica]K83
    |    |--S. curtula Champion 1908D09
    |    |--S. punctipennis Hustache 1951H51
    |    `--S. subplanipennis Hustache 1951 [=Madarellus planipennis ms]H51
    |--MicrostratesD09
    |    |--M. bipunctatus Hustache 1951H51
    |    |--M. cocois Bondar 1941D09
    |    |--M. rufus Hustache 1951H51
    |    `--M. ypsilon Mshl. 1920 [=M. scutellatus Hustache 1951 (n. n.)]H51
    |--Leptoschoinella Hustache 1951H51
    |    |--*L. croceosignata (Boh. 1844) [=Centrinus croceosignatus]H51
    |    |--L. humeralis Hustache 1951H51
    |    |--L. quadripunctata Hustache 1951H51
    |    `--L. sericea Hustache 1951H51
    |--Conoproctus Lacordaire 1866D09, K83 [incl. Leptomadarus Casey 1922K83, Pseudomadarus Casey 1922K83]
    |    |--C. biplagiatus (Boheman 1836) [=Madarus biplagiatus; incl. M. latus Casey 1922]K83
    |    |--C. exaratus (Hustache 1951)K83, H51 [=Madarus (Eumadarus) exaratusK83, H51]
    |    |--C. longipes (Casey 1922) [=Madarus longipes; incl. M. productus Casey 1922]K83
    |    |--C. quadriplagiatus Lacordaire 1866 [incl. C. fluvialis Casey 1922]K83
    |    `--C. quadripustulatus Fabricius 1801D09 (see below for synonymy)
    |--PlatyonyxD09
    |    |--P. circumscriptus Hustache 1951H51
    |    |--P. interruptus Hustache 1951H51
    |    |--P. ornatus Schoenherr 1826D09 (see below for synonymy)
    |    |--P. signatus [=Centrinus signatus]K83
    |    `--P. suturalis [=Leptoschoinus suturalis]K83
    |--Madarus Schönherr 1825 [incl. Eumadarus Casey 1922]K83
    |    |--*M. corvinusK83
    |    |--M. bistrigellus Boheman 1844D09 [=Linomadarus bistrigellusK83]
    |    |--M. impressirostris [=M. (Eumadarus) impressirostris]H51
    |    |--M. nemorosus Hustache 1951 [=M. (Eumadarus) nemorosus]H51
    |    |--M. oblongus Hustache 1951 [=M. (Eumadarus) oblongus]H51
    |    `--M. scutellatus Hustache 1951 [=M. (Eumadarus) scutellatus]H51
    |--Ampeloglypter LeConte 1876A02
    |    |--A. brunnescens Hustache 1951K83, H51 [=A. brunescensK83]
    |    |--A. cayennensis Hustache 1951H51
    |    |--A. heterosternoides Hustache 1951H51
    |    |--A. laevis Hustache 1951H51
    |    |--A. longiclava Hustache 1951H51
    |    |--A. pilosellus Hustache 1951H51
    |    |--A. singularis Hustache 1951H51
    |    `--A. vicinus Hustache 1951H51
    |--Madarellus Casey 1892 [incl. Willinkia Bondar 1949]A02
    |    |--M. castaneus Hustache 1951H51
    |    |--M. claveri Hustache 1951H51
    |    |--M. cuneipennis Hustache 1951H51
    |    |--M. denieri Hustache 1951H51
    |    |--M. ebenus (Fabricius 1801) [incl. M. laticollis Boheman 1844]K83
    |    |--M. ruficollisH51
    |    |--M. sanguinicollis Hustache 1951H51
    |    |--M. singularis (Boheman 1844) [incl. M. metallicus Bondar 1949]K83
    |    `--M. vinaceus Hustache 1951H51
    `--LinomadarusD09
         |--L. bilineatus [=Madarus bilineatus]K83
         |--L. binotatusH51
         |--L. bisignatus [=Madarus bisignatus]K83
         |--L. boliviensis Hustache 1951H51
         |--L. brevilinea [=Madarus brevilinea]K83
         |--L. bridarollii [=Paramadarus bridarollii]K83
         |--L. claveri Hustache 1951H51
         |--L. convexipennis Hustache 1951H51
         |--L. discalis Casey 1922 [incl. L. aeneotinctus Hustache 1940, L. sylvaticus Hustache 1940]K83
         |--L. distigma [=Madarus distigma]K83
         |--L. fulvitarsis Hustache 1951H51
         |--L. macrogrammus [=Madarus macrogrammus]K83
         |--L. monostigma Hustache 1951H51
         |--L. perlaevis Casey 1922 [incl. L. subtilis Casey 1922]K83
         `--L. vorticosus Casey 1922D09

Conoproctus quadripustulatus Fabricius 1801^D09 [=Madarus quadripustulatus^K83; incl. M. barbatulus Casey 1922^K83, M. lucidus Casey 1922^K83, M. parensis Casey 1922^K83, M. vividus Casey 1922^K83]

Platyonyx ornatus Schoenherr 1826^D09 [incl. P. ornatus var. diffusus Hustache 1951^H51, P. ornatus var. griseofasciatus Hustache 1951^H51]

*Type species of generic name indicated

References

[A02] Anderson, R. S. 2002. Curculionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 722–815. CRC Press.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D09] Davis, S. R. 2009. Morphology of Baridinae and related groups (Coleoptera, Curculionidae). ZooKeys 10: 1–136.

[H51] Hustache, A. 1951. Nouveaux Barinae Sud-Américains. 4.^ème partie: Madopterina—Lyterida—Leptoschoina—Eutoxina—Madarina—Barymerina. Boletim do Museu Nacional, nova serie, Zoologia 102: 1–78.

[K83] Kuschel, G. 1983. New synonymies and combinations of Baridinae from the Neotropic and Nearctic regions (Coleoptera: Curculionidae). Coleopterists Bulletin 37 (1): 34–44.

Lacordaire, T. 1866. Histoire Naturelle des Insectes. Genera des Coléoptères, ou exposé méthodique et critique de tous les genres proposés jusqu’ici dans cet ordre d’insectes vol. 7. Librairie encyclopédique de Roret: Paris.

[L18] Legalov, A. A. 2018. Annotated key to weevils of the world. Part 3. Subfamily Conoderinae (Coleoptera, Curculionidae). Ukrainian Journal of Ecology 8 (4): 494–503.