Belongs within: Araneida.
The Mesothelae are a group of spiders in which the abdomen retains external segmentation, the fangs are obliquely angled and the spinnerets are more or less displaced to the centre of the abdomen (Wunderlich 2019). Living members of the group are restricted to eastern Asia and inhabit tubular burrows with a trapdoor (Jocqué & Dippenaar-Schoeman 2007). Fossil representatives include the Arthrolycosidae, known from the Carboniferous and Permian of Eurasia and North America (Fritsch 2004).
<==Mesothelae [Liphistiomorphae, Verticulatae]
| i. s.: Palaeothele montceauensisSSS08
| ‘Eothele’ Selden 2000 non Rowell 1980W19
| Vinathela nahangW19
|--ArthromygalidaeS93
| |--Protocteniza britannica Petrunkevitch 1949S93
| `--ProtolycosaS93
| |--P. anthracophyla Römer 1866F04
| `--P. cebennensis Laurentiaux-Vieira & Laurentiaux 1963S93
|--ArthrolycosidaeS93
| |--Geralycosa fritschii Kušta 1888F04
| |--Rakovnicia antiqua Kušta 1884F04
| |--Eotarbis litoralis Kušta 1888F04
| |--Eocteniza silvicola Pocock 1911S93
| `--Arthrolycosa [incl. Eolycosa Kušta 1885, Scudderia Kušta 1888]F04
| |--A. antiqua Harger 1874S93
| |--A. beecheri Fritsch 1904F04
| |--‘Palaranea’ borassifoliae [=Arthrolycosa palaranea Fritsch 1904]F04
| |--A. carbonaria (Kušta 1888) [=*Scudderia carbonaria]F04
| |--A. danielsi Petrunkevitch 1913S93
| |--A. fortis Fritsch 1904F04
| `--A. lorenzi (Kušta 1885) [=*Eolycosa lorenzi]F04
|--EomesothelidaeW19
| |--Eomesothele Wunderlich 2019W19
| | `--*E. noninclinata Wunderlich 2019W19
| `--Intermesothele Wunderlich 2019W19
| `--*I. pulcher Wunderlich 2019W19
|--Burmathele Wunderlich 2017 [Burmathelidae]W19
| `--B. biseriata Wunderlich 2017W19
|--Cretaceothele [Cretaceothelidae]W19
| `--C. lata Wunderlich 2015W19
|--ParvithelidaeW19
| |--Parvithele Wunderlich 2017W19
| `--Pulvillothele haupti Wunderlich 2017W19
`--LiphistiidaeJD-S07
| i. s.: Nanthela Haupt 2003JD-S07
| Songthela Ono 2000JD-S07
| Palaeocteniza crassipesB86
|--+--Ryuthela Haupt 1983WC17, JD-S07
| | `--R. nishihirai (Haupt 1969)H07
| `--Heptathele Kishida 1923WC17, JD-S07 [Heptathelidae]
| |--H. hangzhouensisWT06
| |--H. nishiharaiH95
| `--H. kimuraiFF99
`--Liphistius Schiödte 1849JD-S07
| i. s.: L. batuensisR95
| L. bristoweiW96
| L. desultorJD-S07
| L. lannaianusJD-S07
| L. malayanusWY88
| L. sumatranusJD-S07
|--L. erawanWC17
`--+--L. bicoloripes (Ono 1988)WC17, GE02
`--L. murphyorumWC17
*Type species of generic name indicated
References
[B86] Bernini, F. 1986. Current ideas on the phylogeny and the adaptive radiations of Acarida. Bolletino di Zoologia 53: 279–313.
[FF99] Forster, R., & L. Forster. 1999. Spiders of New Zealand and their World-wide Kin. University of Otago Press: Dunedin (New Zealand).
[F04] Fritsch, A. 1904. Palaeozoische Arachniden. Selestverlag: Prague.
[GE02] Giribet, G., G. D. Edgecombe, W. C. Wheeler & C. Babbitt. 2002. Phylogeny and systematic position of Opiliones: a combined analysis of chelicerate relationships using morphological and molecular data. Cladistics 18: 5–70.
[H07] Haupt, J. 2007. The whipscorpion Minbosius manilanus (C. L. Koch 1843) on Panay Island (Philippines) (Arachnida, Uropygi, Thelyphonidae). Senckenbergiana Biologica 87 (2): 135–136.
[H95] Humphreys, W. F. 1995. Chorion surface features of chelicerate eggs. Records of the Western Australian Museum Supplement 52: 171–181.
[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).
[R95] Raven, R. J. 1995. Coxal glands of spiders of the genera Bymainiella, Atrax and Namea (Hexathelidae, Dipluridae, Mygalomorphae). Records of the Western Australian Museum Supplement 52: 67–71.
[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.
[SSS08] Selden, P. A., W. A. Shear & M. D. Sutton. 2008. Fossil evidence for the origin of spider spinnerets, and a proposed arachnid order. Proceedings of the National Academy of Sciences of the USA 105 (52): 20781–20785.
[WT06] Waeschenbach, A., M. J. Telford, J. S. Porter & D. T. J. Littlewood. 2006. The complete mitochondrial genome of Flustrellidra hispida and the phylogenetic position of Bryozoa among the Metazoa. Molecular Phylogenetics and Evolution 40 (1): 195–207.
[WY88] Welbourn, W. C., & O. P. Young. 1988. Mites parasitic on spiders, with a description of a new species of Eutrombidium (Acari, Eutrombidiidae). Journal of Arachnology 16: 373–385.
[W96] Wheeler, W. 1996. Optimization alignment: the end of multiple sequence alignment in phylogenetics? Cladistics 12: 1–9.
[WC17] Wheeler, W. C., J. A. Coddington, L. M. Crowley, D. Dimitrov, P. A. Goloboff, C. E. Griswold, G. Hormiga, L. Prendini, M. J. Ramírez, P. Sierwald, L. Almeida-Silva, F. Alvarez-Padilla, M. A. Arnedo, L. R. Benavides Silva, S. P. Benjamin, J. E. Bond, C. J. Grismado, E. Hasan, M. Hedin, M. A. Izquierdo, F. M. Labarque, J. Ledford, L. Lopardo, W. P. Maddison, J. A. Miller, L. N. Piacentini, N. I. Platnick, D. Polotow, D. Silva-Dávila, N. Scharff, T. Szűts, D. Ubick, C. J. Vink, H. M. Wood & J. Zhang. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33: 574–616.
[W19] Wunderlich, J. 2019. What is a spider? Cretaceous fossils modify strongly phylogenetics as well as diagnoses of families, superfamilies and even suborders of spiders (Araneida) and other arthropods. Beiträge zur Araneologie 12: 1–32.