Myrtales

Belongs within: Malvidae.
Contains: Combretaceae, Onagraceae, Lythraceae, Melastomatineae, Myrtaceae.

The Myrtales are a clade of flowering plants, most members of which have flaky bark and opposite leaves with colleters (Angiosperm Phylogeny Website).

Synapomorphies (from Angiosperm Phylogeny Website): Bark flaky; flavonols only, myricetin, methylated ellagic acid present; cork cambium deep seated; pits vestured; libriform fibres septate; tension wood present; secondary phloem stratified; vascular bundles bicollateral, internal phloem present; nodes 1:1; cuticle waxes often absent; branching from current flush; leaves opposite, lamina with brochidodromous venation, colleters present; inflorescence racemose; hypanthium present, nectariferous; calyx valvate, corolla clawed; anthers incurved in bud; pollen with pseudocolpi; gynoecium inferior, style long, minor stylar bundles present, stigma wet; ovules many/carpel, micropyle bistomal and zig-zag, inner integument ca 2 cells across; antipodal cells ephemeral; exotegmen cells tracheidal, endotesta crystalliferous; endosperm at most slight.

<==Myrtales [Myrtanae, Sonneratiaceae]
    |  i. s.: XenodendronYY22
    |         PaleomyrtinaeaWM09
    |         Sonneratiorhizos raoii Chitaley 1969CBH93
    |         SonneratioxylonCBH93
    |         Florschuetzia levipoliCBH93
    |--Lythrineae [Onagrineae]T00
    |    |--CombretaceaeWM09
    |    `--+--OnagraceaeWM09
    |       `--LythraceaeWM09
    `--+--MelastomatineaeWM09
       `--MyrtineaeT00
            |--MyrtaceaeWM09
            `--VochysiaceaeWM09
                 |--Qualeoxylon itaquaquecetubense Suguio & Mussa 1978CBH93
                 `--QualeaWM09
                      |--Q. grandifloraWO05
                      `--Q. parvifloraWM09

*Type species of generic name indicated

References

[CBH93] Collinson, M. E., M. C. Boulter & P. L. Holmes. 1993. Magnoliophyta (‘Angiospermae’). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 809–841. Chapman & Hall: London.

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441–647.

[WM09] Wang, H., M. J. Moore, P. S. Soltis, C. D. Bell, S. F. Brockington, R. Alexandre, C. C. Davis, M. Latvis, S. R. Manchester & D. E. Soltis. 2009. Rosid radiation and the rapid rise of angiosperm-dominated forests. Proceedings of the National Academy of Sciences of the USA 106 (10): 3853–3858.

[WO05] Wilson, P. G., M. M. O’Brien, M. M. Heslewood & C. J. Quinn. 2005. Relationships within Myrtaceae sensu lato based on a matK phylogeny. Plant Systematics and Evolution 251: 3–19.

[YY22] Yampolsky, C., & H. Yampolsky. 1922. Distribution of sex forms in the phanerogamic flora. Bibliotheca Genetica 3: 1–62.