Neonemertea

Light-margined ribbon worm Cerebratulus marginatus, photographed by James Zhan.

Belongs within: Nemertea.
Contains: Heteronemertea, Monostilifera.

The Neonemertea is a clade of nemertean worms identified by molecular analysis (Thollesson & Norenburg 2003). It unites the clades Hoplonemertea, in which the eversible proboscis is armed by one or more stylets, and Pilidiophora, whose members possess a distinctive larval form called a pilidium.

<==Neonemertea
    |--PilidiophoraTN03
    |    |--HeteronemerteaTN03
    |    |--HubrechtidaeTN03
    |    |    |--HubrechtiaTN03
    |    |    `--Sundbergia Gibson 2002G02
    |    |         `--*S. albula Gibson 2002G02
    |    `--HubrechtellidaeC03a
    |         |--TetramysC03a
    |         |--CoeiaC03a
    |         `--Hubrechtella Bergendal 1902C03a
    |              |--H. alba Gibson 1997C03a
    |              |--H. dubia Bergendal 1902TN03
    |              |--H. juliae Chernyshev 2003C03a
    |              |--H. malabarensis Gibson 1979C03a
    |              `--H. sinimarinus Gibson & Sundberg 1999C03a
    `--Hoplonemertea [Enopla, Herponemertoidea, Hoplonemertini, Metanemertini, Polystilifera]TN03
         |  i. s.: ArenonemertesK86
         |           |--A. microps Friedrich 1933K86
         |           `--A. minutus Friedrich 1949K86
         |         *Prostomatella arenicola Friedrich 1935K86
         |         OtonemertesK86
         |           |--O. denisi Dawydoff 1937K86
         |           `--O. marcusi Corrêa 1958K86
         |         GeonemertesG02
         |           |--G. caeca Darbishire 1909 (n. d.)G02
         |           |--G. pelaensisM06
         |           `--G. spirospermia Darbishire 1909 (n. d.)G02
         |         Crassonemertes robustaC03b
         |         Tortus tokmakovaeC03b
         |         Collarenemertes bimaculataC03b
         |         GononemertesC03b
         |--PelagicaC03b
         |    |  i. s.: PlanktonemertesC03b
         |    |--Nectonemertes mirabilis Verrill 1892TN03
         |    `--+--PelagonemertesTN03
         |       `--+--Phallonemertes murrayi (Brinkmann 1912)TN03
         |          `--ProtopelagonemertesTN03
         `--+--MonostiliferaTN03
            `--ReptantiaC03b
                 |--Siboganemertes [Archireptantia, Siboganemertidae]C03b
                 |    `--S. weberiC03b
                 `--EureptantiaC03b
                      |--Sagaminemertes Friedrich 1968 [Sagaminemertida, Sagaminemertidae]C03b
                      `--+--Uniporidae [Uniporida]C03b
                         `--Drepanophorida [Tetraophthalmida]C03b

*Type species of generic name indicated

References

[C03a] Chernyshev, A. V. 2003a. New species of the genus Hubrechtella (Nemertea, Anopla) from the Sea of Japan and validation of the family Hubrechtellidae. Russian Journal of Marine Biology 29: 333–336.

[C03b] Chernyshev, A. V. 2003b. Classification system of the higher taxa of enoplan nemerteans (Nemertea, Enopla). Biologiya Mariya—Marine Biology 29: 299–306 (translated: Russian Journal of Marine Biology 29 (Suppl. 1): S57–S65).

[G02] Gibson, R. 2002. The Invertebrate Fauna of New Zealand: Nemertea (ribbon worms). NIWA Biodiversity Memoir 118. National Institute of Water and Atmospheric Research: Wellington.

[K86] Kirsteuer, E. 1986. Nemertina. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 72–75. E. J. Brill/Dr W. Backhuys: Leiden.

[M06] Moore, J. 2006. An Introduction to the Invertebrates 2nd ed. Cambridge University Press.

[TN03] Thollesson, M., & J. L. Norenburg. 2003. Ribbon worm relationships: a phylogeny of the phylum Nemertea. Proceedings of the Royal Society of London Series BBiological Sciences 270: 407–415.

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