Belongs within: Neritoidea.
Contains: Smaragdia, Nerita.
The Neritidae are marine gastropods characterised by a shell with dissolved inner whorls and radula with large, smooth, mineralised lateral teeth with a cutting and scraping edge (Bandel 2001). The Eocene Velates were limpet-like neritids with the base of the shell formed by the callus of the inner lip (Bandel 2001).
Neritidae [Neritae, Neritinia]
|--Velates Montfort 1810 [Velatinae]BR05
| |--V. schmiedelianus Chemnitz 1886B01 (see below for synonymy)
| |--V. balkanicus Bontscheff 1897B01
| |--V. californicus Vokes 1935B01
| |--V. noetlingi Cossmann & Pisarro 1909B01
| |--V. perversus (Gmelin 1791)KC60 [=Nerita perversaB01]
| `--V. vizcainoensis Cushing-Woods & Saul 1986B01
|--Smaragdiinae [Smaragdiidae, Smaragdinae]BR05
| |--SmaragdiaBR05
| |--Tanzaniella Lupu 1979B01
| | `--*T. souverbiana (Montrozier 1883)B01 [=Neritina souverbianaB01; incl. N. (Vitta) pulcherrima Angas 1871PB27]
| |--Neodesmieria Magne & Vergneau-Saubade 1974B01
| | `--*N. sanctistephani (Cossmann & Peyrot 1917) [=Nerita sanctistephani]B01
| `--Magadis Melvill & Standen 1899KC60
| `--*M. eumerintha Melvill & Standen 1899KC60
`--Neritinae [Protoneritidae]BR05
|--NeritaBR05
|--Protonerita Kittl 1894BR05
| `--*P. calcitica Kittl 1894BR17 [=Neritaria calciticaTE01]
|--Semineritina Cossmann 1925 [=Seminerita Wenz 1938]B01
| `--*S. mammaria (Lamarck 1804)B01 [=Nerita mammariaKC60]
|--Bathynerita Clarke 1989B01
| `--*B. naticoidea Clarke 1989B01
|--Cryptonerita Kittl 1894 [=Cryptonatica (l. c.) non Dall 1892]KC60
| `--*C. elliptica Kittle 1894KC60
|--Trochonerita Cossmann 1907KC60
| `--*T. mammaeformis (d’Orbigny 1850) [=Nerita mammaeformis]KC60
|--Tournouerella Munier-Chalmas 1887 [=Tournoueria Munier-Chalmas 1884 non Brusina 1870]KC60
| `--*T. matheroni (Munier-Chalmas 1884) [=*Tournoueria matheroni]KC60
|--Trachynerita Kittl 1894B01
| |--*T. fornoensis Kittl 1894TE01
| |--T. depressaKC60
| `--T. quadrata (Stoppani 1858) [=Turbo quadrata]KC60
|--Mesoneritina Yen 1946B01
| |--*M. nebrascensis (Meek & Hayden 1862)B01 [=Neritella nebrascensisBK03]
| |--M. ajkaensis Bandel & Kiel 2003BK03
| |--M. liasinaG88
| |--M. morrisonensisG88
| `--M. subhercynica (Mertin 1939)BK03
|--Neritoma Morris 1849KC60
| | i. s.: N. nudaB01
| | N. woodfordi Allison 1955B01
| |--*N. (Neritoma) sinuosa (Sowerby 1818)B01 [=Nerita sinuosaKC60]
| `--N. (Staffinia Anderson & Cox 1948)KC60
| `--N. (*S.) staffinensis (Forbes 1851) [=Neritina staffinensis]KC60
|--Lyosoma White 1880KC60, B01
| |--*L. powelli White 1876B01
| |--‘Nerita’ costiferaB01
| |--‘Nerita’ costulata Deshayes 1838B01
| |--‘Nerita’ pseudocostata Orbigny 1849B01
| `--‘Nerita’ subrugosa Hudleston 1894B01
|--Puperita Gray 1857B01
| |--*P. pupa (Linné 1767)B01 [=Nerita pupaKC60, Neritina pupaB01]
| |--P. bensoni (Récluz 1850)B01
| |--P. japonicaB01
| |--P. tristis (Orbigny 1842)B01
| `--P. vincta Usticke 1969BC01
`--Neritoplica Oppenheim 1892KC60 [incl. Bajanerita Squires 1993BK03]
|--*N. uniplicata (Sowerby 1823)BK03 (see below for synonymy)
|--N. dockeryi Bandel & Kiel 2003BK03
|--N. loganensis (Erickson 1974) [=Neritina loganensis]BK03
|--N. matheroni Oppenheim 1895BK03
|--N. oppenheimi Bandel & Kiel 2003BK03
|--N. pisum (Meek 1873)BK03
|--N. subornata (Orbigny 1850)B01
`--N. trochispira Bandel & Kiel 2003BK03
Neritidae incertae sedis:
Agapilia Harzhauser & Kowalke 2001GK02, B01
`--*A. picta (Férussac 1825) [=Nerita picta, Neritina picta]B01
Tomostoma Deshayes 1823B01 [incl. Calana Gray 1847KC60]
|--*T. neritoides (Deshayes 1823)B01 [=Pileolus neritoidesKC60]
`--T. dudariensis (Strausz 1966) [=Pileolus (Tomostoma) dudariensis]B01
Neritoptyx Oppenheim 1892BK03
|--*N. goldfussi (Keferstein 1829) [=Nerita goldfussi, Neritopsis goldfussi]BK03
|--N. renauxiana (d’Orbigny 1842)BK03
`--N. reticulirata (Dockery 1993) [=Nerita reticulirata]BK03
Schwardtina Bandel & Riedel 1994BK03
`--*S. cretacea (Tausch 1886) [=Helix cretacea]B01
*Neritoplica uniplicata (Sowerby 1823)BK03 [=Neritina uniplicataBK03; incl. Nerita globosa Ferussac 1823KC60, Neritina globulus Deshayes 1824BK03, Neritoplica globulusB01]
Velates schmiedelianus Chemnitz 1886B01 [=Neritina schmiedelianaB01; incl. *V. conoideus Montfort 1810BR17, N. conoideaB01]
*Type species of generic name indicated
References
[B01] Bandel, K. 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen des Geologisch-Paläontologischen Institutes der Universität Hamburg 85: 65–164.
[BK03] Bandel, K., & S. Kiel. 2003. Relationships of Cretaceous Neritimorpha (Gastropoda, Mollusca), with the description of seven new species. Bulletin of the Czech Geological Survey 78 (1): 53–65.
[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.
[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.
[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.
[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.
[GK02] Gründel, J., & T. Kowalke. 2002. Palaeorissoinidae, a new family of marine and brackish water Rissooidea (Gastropoda, Littorinimorpha). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 226 (1): 43–57.
[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.
[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.
[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1–47.