Belongs within: Ocenebrinae.
Nucella is a genus of muricid snails with an acuminate spire, foliated or tuberculose whorls, an anteriorly narrowed aperture, and small and oblique siphonal canal.
Nucella Röding 1798BR05 [incl. Polytropa Swainson 1840RD75; Nucellidae]
| i. s.: ‘Buccinum’ filosum Gmelin 1791 [=*Nucella theobroma Röding 1798]BR17
| N. tokudaiAV03
|--N. lapillus (Linnaeus 1758)RF23, RD75 [=Buccinum lapillusRD75, *Polytropa lapillusRD75, Purpura lapillusD56]
`--+--N. canaliculata (Duclos 1832)RF23, O27 [=Thais (Nucella) canaliculataO27]
| |--N. c. canaliculataO27
| |--‘Thais’ c. analoga Forbes 1850O27
| `--‘Thais’ c. compressa Dall 1915O27
|--N. heyseanaRF23
|--N. lamellosa (Gmelin 1792)RF23, O27 [=Thais (Nucella) lamellosaO27]
| |--N. l. lamellosaO27
| |--‘Thais’ l. cymica Dall 1915O27
| |--‘Thais’ l. franciscana Dall 1915O27
| |--‘Thais’ l. hormica Dall 1915O27
| |--‘Thais’ l. neptunea Dall 1915O27
| `--‘Thais’ l. sitkana Dall 1915O27
|--+--N. freycinetiRF23
| | |--N. f. freycinetiAV03
| | `--N. f. saitoiAV03
| `--N. lima (Martyn 1784)RF23, O27 [=Thais (Nucella) limaO27]
`--+--N. emarginata (Deshayes 1839)RF23, O27 [=Thais (Nucella) emarginataO27]
| |--N. e. emarginataO27
| `--‘Thais’ e. projecta Dall 1915O27
`--N. ostrina (Gould 1852)RF23, O27 [=Thais emarginata ostrinaRF23]
*Type species of generic name indicated
References
[AV03] Amano, K., & G. J. Vermeij. 2003. Evolutionary adaptation and geographic spread of the Cenozoic buccinid genus Lirabuccinum in the North Pacific. Journal of Paleontology 77 (5): 863–872.
[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.
[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.
[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms 2nd ed. University Press: Cambridge.
[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 2. Stanford University Press: Stanford University (California).
[RD75] Radwin, G. E., & A. D’Attilio. 1975. A catalogue of muricacean generic taxa. Transactions of the San Diego Society of Natural History 17 (20): 279–292.
[RF23] Russini, V., G. Fassio, E. Nocella, R. Houart, A. Barco, N. Puillandre, P. Lozouet, M. V. Modica & M. Oliverio. 2023. Whelks, rock-snails, and allied: a new phylogenetic framework for the family Muricidae (Mollusca: Gastropoda). European Zoological Journal 90 (2): 856–868.