
Belongs within: Linguloidea.
The Obolinae are an assemblage of inarticulate brachiopods known from the Lower Cambrian to the Lower Carboniferous (Holmer & Popov 2000).
Characters (Holmer & Popov 2000): Shell rounded, elongate, suboval to subtriangular, unequivalved; ventral pseudointerarea well developed, raised or continuous with valve floor; dorsal pseudointerarea with median groove or undivided.
<==Obolinae [Lingulellidae, Lingulellinae]HP00
|--Pseudobolus Cooper 1956R65
| `--*P. gibbosus (Willard 1928) [=Palaeoglossa gibbosa]R65
|--Schmidtites Schuchert & LeVene 1929 (see below for synonymy)R65
| `--*S. celatus (Volborth 1869) [=*Schmidtia celata]R65
|--Lingulepis Hall 1863R65
| |--*L. pinnaformis (Owen 1852) [=Lingula pinnaformis]R65
| |--L. acuminata (Conrad 1839) [=Lingula (Glossina) acuminata, Lingulella (Lingulepis) acuminata]W08
| | |--L. a. acuminataW08
| | `--L. a. sequens (Walcott 1908) [=Lingulella (Lingulepis) acuminata sequens]W08
| `--L. longinervis (Matthew 1903) [=Lingulella (Lingulepis) longinervis]W08
|--Spinilingula Cooper 1956R65
| `--*S. intralamellata Cooper 1956R65
|--Agalatassia Popov & Holmer 1994HP00
| `--*A. triangularis Popov & Holmer 1994HP00
|--Aksarinaia Koneva 1992HP00
| `--*A. triquetra Koneva 1992HP00
|--Anomaloglossa Percival 1978HP00
| `--*A. porca Percival 1978HP00
|--Barbatulella Williams & Lockley 1983HP00
| `--*B. lacunosa Williams & Lockley 1983HP00
|--Chakassilingula Ushatinskaya 1992HP00
| `--*C. erbiensis Ushatinskaya 1992HP00
|--Euobolus Mickwitz 1896HP00
| `--*E. elegans (Mickwitz 1896) [=Obolus (*Euobolus) elegans]HP00
|--Expellobolus Havlíček 1982HP00
| `--*E. expulsus (Barrande 1879) [=Lingula expulsa]HP00
|--Experilingula Koneva & Popov 1983HP00
| `--*E. divulgata Koneva & Popov 1983HP00
|--Fordinia Walcott 1908HP00
| |--*F. perfecta (Walcott 1908) [=Obolus (*Fordinia) perfecta]HP00
| |--F. bellula (Walcott 1905) [=Obolus (Fordinia) bellulus]W08
| `--F. gilberti (Walcott 1908) [=Obolus (Fordinia) gilberti]W08
|--Foveola Gorjansky 1969 [=Faveolla Popov in Nazarov & Popov 1980]HP00
| `--*F. maarduensis Gorjansky 1969HP00
|--Gorjanskya Tenjakova 1980HP00
| `--*G. tselinogradica Tenjakova 1980HP00
|--Leontiella Yadrenkina 1982 [=Khalfinella Yadrenkina 1982]HP00
| `--*L. gloriosa Yadrenkina 1982 [=*Khalfinella gloriosa]HP00
|--Leptembolon Mickwitz 1896HP00
| `--*L. lingulaeformis (Mickwitz 1896) [=Obolus (*Leptembolon) lingulaeformis]HP00
|--Lindinella Mergl & Šlehoferová 1990HP00
| `--*L. kordulei Mergl & Šlehoferová 1990HP00
|--Notiobolus Popov in Popov & Soloviev 1981HP00
| `--*N. tenuis Popov in Popov & Soloviev 1981HP00
|--Palaeoglossa Cockerell 1911 [=Glossina Phillips 1848 non Wiedemann 1830, Palaeoglossina Dunbar & Condra 1932]HP00
| `--*P. attenuata (Sowerby 1839) [=Lingula attenuata, *Glossina attenuata]R65
|--Palaeoschmidtites Koneva 1979HP00
| `--*P. minimus Koneva 1979HP00
|--Ralfia Popov & Khazanovitch in Popov et al. 1989HP00
| `--*R. ovata (Pander 1830) [=Ungula ovata]HP00
|--Rebrovia Popov & Khazanovitch in Popov et al. 1989HP00
| `--*R. chernetskae Popov & Khazanovitch in Popov et al. 1989HP00
|--Sinoglossa Mei 1993HP00
| `--*S. amygdalina Mei 1993HP00
|--Squamilingulella Xu 1978HP00
| `--*S. zhonghuaensis Xu 1978HP00
|--Westonisca Havlíček 1982HP00
| `--*W. lamellosa (Barrande 1879) [=Lingula lamellosa]HP00
|--Dicellomus Hall 1871R65
| |--*D. politus (Hall 1861) [=Lingula polita]R65
| |--D. occidentalisR65
| |--D. parvus Walcott 1908W08
| `--D. prolificus Walcott 1908W08
|--Lingulobolus Matthew 1895 [incl. Sphaerobolus Matthew 1895 non Tode 1790 (ICBN)]R65
| |--*L. affinis (Billings 1872) [=Lingulella affinis]R65
| `--L. spissus (Billings 1872)HP00, R65 [=Lingulella spissaR65, *Sphaerobolus spissusR65]
|--Obolus von Eichwald 1829 [=Obulus Quenstedt 1868]HP00
| |--*O. apollinis von Eichwald 1829HP00 [=Euobolus apollinisR65]
| |--O. discoideus (Hall & Whitfield 1877)W08
| |--O. feistmanteli (Barrande 1879)W08
| |--O. lamborni (Meek 1871)W08
| |--O. membranaceous Walcott 1908W08
| |--O. minimus Walcott 1905W08
| |--O. mootwingensis Fletcher 1964F71
| |--O. parvus Walcott 1908W08
| |--O. smithi Walcott 1908W08
| |--O. tetonensis Walcott 1901W08
| | |--O. t. tetonensisW08
| | `--O. t. leda Walcott 1908W08
| |--O. willisi (Walcott 1898)W08
| `--O. wortheni Walcott 1908W08
|--Westonia Walcott 1901R65
| |--*W. aurora (Hall 1861) [=Lingula aurora]R65
| |--W. bottnica (Wiman 1902) [=Obolus (Westonia) bottnica]W08
| |--W. dartoni (Walcott 1908) [=Obolus (Westonia) dartoni]W08
| |--W. ella (Hall & Whitfield 1877) [=Obolus (Westonia) ella]W08
| | |--W. e. ellaW08
| | `--W. e. onaquiensis (Walcott 1908) [=Obolus (Westonia) onaquiensis]W08
| |--W. elongatus (Walcott 1908) [=Obolus (Westonia) elongatus]W08
| |--W. escasoni (Matthew 1901) [=Obolus (Westonia) escasoni]W08
| |--W. euglyphus (Walcott 1908) [=Obolus (Westonia) euglyphus]W08
| |--W. finlandensis (Walcott 1902) [=Obolus (Westonia) finlandensis]W08
| |--W. iphis (Walcott 1905) [=Obolus (Westonia) iphis]W08
| |--W. linguloidesR65
| |--W. notchensis (Walcott 1908) [=Obolus (Westonia) notchensis]W08
| |--W. stoneanus (Whitfield 1882) [=Obolus (Westonia) stoneanus]W08
| `--W. wasatchensis (Walcott 1908) [=Obolus (Westonia) wasatchensis]W08
|--Apatobolus Popov in Nazarov & Popov 1980 [incl. Paldiskites Havlíček 1982]HP00
| |--*A. plicatus Popov in Nazarov & Popov 1980HP00
| `--‘Lingula’ sulcata Barrande 1879 [=*Paldiskites sulcata]HP00
|--Kyrshabaktella Koneva 1986 [=Kyrshabactella Ushatinskaya 1992]HP00
| |--*K. certa Koneva 1986HP00
| `--K. belli (Pelman 1977)HB93
|--Oepikites Khazanovitch & Popov in Khazanovich, Popov & Melnikova 1984HP00
| |--*O. macilentus Khazanovitch & Popov in Khazanovich, Popov & Melnikova 1984HP00
| `--O. triquetrusHP00
|--Ungula Pander 1830 [incl. Ungulites Quenstedt 1837]HP00
| |--*U. convexa Pander 1830R65
| `--U. ingricaW97
|--Palaeobolus Matthew 1899R65 [=Palaebolus (l. c.)R65; incl. Obolopsis Saito 1936HP00]
| |--*P. bretonensis Matthew 1899R65
| |--P. liantuoensisL04
| `--‘*Obolopsis’ margaritae Saito 1936HP00
|--Trigonoglossa Dunbar & Condra 1932HP00
| |--*T. nebrascensis (Meek 1872) [=Lingula nebrascensis]R65
| |--T. scotica (Davidson 1860)HB93
| `--T. spatiosaHP00
`--Lingulella Salter 1866R65
|--*L. davisii (M’Coy 1851) [=Lingula davisii]R65
|--L. ampla (Owen 1852)W08
|--L. antiquissimaHP00
|--L. buttsi Walcott 1908W08
|--L. chengjiangensisZH04
|--L. ferrugineaT10
|--L. gnaltaensis Fletcher 1964 [=L. (Leptembolon) gnaltaensis]F71
|--L. incurvataLM05
|--L. manticula (White 1874)W08
|--L. texana Walcott 1908W08
`--L. variabilis Pelman 1977HB93
Inorganic?: Lingulella montana Fenton & Fenton 1936H75
Schmidtites Schuchert & LeVene 1929 [=Schmidtia Volborth 1869 nec Steud. ex Schmidt 1852 (ICBN) nec Balsamo-Crivelli 1863]R65
*Type species of generic name indicated
References
[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.
[H75] Häntzschel, W. 1975. Treatise on Invertebrate Paleontology pt W. Miscellanea. Supplement 1. Trace Fossils and Problematica, 2nd ed. The Geological Society of America: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).
[HB93] Harper, D. A. T., C. H. C. Brunton, L. R. M. Cocks, P. Copper, E. N. Doyle, A. L. Jeffrey, E. F. Owen, M. A. Parkes, L. E. Popov & C. D. Prosser. 1993. Brachiopoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 427–462. Chapman & Hall: London.
[HP00] Holmer, L. E., & L. E. Popov. 2000. Lingulata. In: Kaesler, R. L. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda, Revised vol. 2. Linguliformea, Craniiformea, and Rhynchonelliformea (part) pp. 30–146. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).
[LM05] Lehnert, O., J. F. Miller, S. A. Leslie, J. E. Repetski & R. L. Ethington. 2005. Cambro-Ordovician sea-level fluctuations and sequence boundaries: the missing record and the evolution of new taxa. Special Papers in Palaeontology 73: 117–134.
[L04] Li G.-X. 2004. Early Cambrian hyolithelminths—Torellella bisulcata sp. nov. from Zhenba, southern Shaanxi. Acta Palaeontologica Sinica 43 (4): 571–578.
[R65] Rowell, A. J. 1965. Inarticulata. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda vol. 1 pp. H260–H296. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas Press: Lawrence (Kansas).
[T10] Taylor, T. G. 1910. The Archaeocyathinae from the Cambrian of South Australia with an account of the morphology and affinities of the whole class. Memoirs of the Royal Society of South Australia 2 (2): 55–188.
[W08] Walcott, C. D. 1908. Cambrian geology and paleontology. No. 3.—Cambrian Brachiopoda: descriptions of new genera and species. Smithsonian Miscellaneous Collections 53: 53–137.
[W97] Williams, A. 1997. Shell structure. In: Kaesler, R. L. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda, Revised vol. 1. Introduction pp. 267–320. The Geological Society of America: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).
[ZH04] Zhang, Z., J. Han, X. Zhang, J. Liu & D. Shu. 2004. Soft-tissue preservation in the Lower Cambrian linguloid brachiopod from South China. Acta Palaeontologica Polonica 49: 259–266.