
Belongs within: Gnathophiuridea.
The Ophiotrichidae are a family of stout-toothed brittle stars with quadrangular, spiniform tooth papillae clustered at the apex of each jaw, and no oral papillae. Their fossil record is limited with questionable exemplars from the Lower Jurassic (Spencer & Wright 1966).
<==Ophiotrichidae
|--Ophiomaza Lyman 1871 [incl. Luetkenia Brock 1888 nec Claus 1864 nec Duncan 1878]SW66
| `--*O. cacaotica Lyman 1871SW66
|--Ophiopteron Ludwig 1888SW66
| `--*O. elegans Ludwig 1888SW66
|--Amphiophiothrix Clark 1946SW66
| `--*A. demessa (Lyman 1861) [=Ophiothrix demessa]SW66
|--Gymnolophus Brock 1888SW66
| `--*G. holdsworthi (Smith 1878) [=Ophiothela holdsworthi]SW66
|--Lissophiothrix Clark 1938SW66
| `--*L. delicata Clark 1938SW66
|--Ophioaethiops Brock 1888 [incl. Ophiohelix Koehler 1895]SW66
| `--*O. unicolor Brock 1888SW66
|--Ophiocnemis Müller & Troschel 1842SW66
| `--*O. marmorata (Lamarck 1816) [=Ophiura marmorata]SW66
|--Ophiolophus Marktanner-Turneretscher 1887SW66
| `--*O. novarae Marktanner-Turneretscher 1887SW66
|--Ophiopeltis Düben & Koren 1846SW66
| `--*O. securigera Düben & Koren 1846SW66
|--Ophiophthirius Döderlein 1898SW66
| `--*O. actinometrae Döderlein 1898SW66
|--Ophiopsammium Lyman 1874SW66
| `--*O. semperi Lyman 1874SW66
|--Ophiosphaera Brock 1888SW66
| `--*O. insignis Brock 1888SW66
|--Ophiotrichoides Ludwig 1882SW66
| `--*O. lymani Ludwig 1882SW66
|--Ophiothela Verrill 1867 [incl. Ophioteresis Bell 1892]SW66
| |--*O. mirabilis Verrill 1867SW66
| `--O. danae Verrill 1869MG-H11
|--Ophiogymna Ljungman 1866 [incl. Ophiocampsis Duncan 1887]SW66
| |--*O. elegans Ljungman 1866SW66
| |--O. pellicula (Duncan 1887)BM09
| `--O. pulchella (Koehler 1905)MG-H11
|--Macrophiothrix Clark 1938 [incl. Placophiothrix Clark 1938]SW66
| |--*M. longipeda (Lamarck 1816) [=Ophiura longipeda]SW66
| |--M. caenosa Hoggett 2006SM15
| |--M. callizona Clark 1938SM15
| |--M. demessa (Lyman 1861)BM09
| |--M. koehleri Clark 1968MG-H11
| |--M. leucosticha Hoggett 1991SM15
| |--M. lorioli Clark 1968SM15
| |--M. megapoma Clark 1938SM15
| |--M. microplax (Bell 1884)SM15
| |--M. paucispina Hoggett 1991SM15
| |--M. propinqua (Lyman 1861)SM15
| |--M. rhabdota (Clark 1915)SM15
| |--M. robillardi (de Loriol 1893)SM15
| |--‘Ophiothrix (Placophiothrix)’ striolata Grube 1868SM15
| `--M. variabilis (Duncan 1887)MG-H11
`--Ophiothrix Müller & Troschel 1840B01 [incl. Ophionyx Müller & Troschel 1840SW66]
|--*O. fragilis (Abildgaard in Müller 1789)TS16 [incl. O. alopecurusBK77, O. echinataBK77, O. versicolorBK77]
|--O. aristulata Lyman 1879MG-H11
|--O. armata Koehler 1905BM09
|--O. caespitosa Lyman 1879MG-H11
|--O. capillarisH04
|--O. ciliaris (Lamarck 1816)MG-H11
|--O. contenta Koehler 1930SM15
|--O. crassispinaW93
|--O. deceptor Koehler 1922SM15
|--O. exigua Lyman 1874BM09
|--O. foveolata Marktanner-Turneretscher 1887MG-H11
|--O. lineocaerulea Clark 1928SM15
|--O. martensi Lyman 1874SM15
|--O. melanosticta Grube 1868SM15
|--O. miles Koehler 1905SM15
|--O. nereidina (Lamarck 1816)BM09
|--O. pentaphylla (Pennant 1777) [=Asterias pentaphylla; incl. Ophiura rosula Fleming 1828, Ophiothrix rosula]SW66
|--O. picteti de Loriol 1893SM15
|--O. plana Lyman 1874SM15
|--O. purpurea von Martins 1867BM09
|--O. quinquemaculataBK77
|--O. royeri (de Loriol 1872)SG93
|--O. savignyi (Müller & Troschel 1842)SM15
|--O. smaragdina Studer 1882SM15
|--O. spiculataCH97
|--O. spongicola Stimpson 1855MG-H11
|--O. trilineata Lütken 1869SM15
`--O. viridialba von Martens 1867MG-H11
*Type species of generic name indicated
References
[BK77] Barel, C. D. N., & P. G. N. Kramers. 1977. A survey of the echinoderm associates of the north-east Atlantic area. Zoologische Verhandelingen 156: 1–159.
[B01] Boczarowski, A. 2001. Isolated sclerites of Devonian non-pelmatozoan echinoderms. Palaeontologia Polonica 59: 1–219.
[BM09] Bryce, C., & L. Marsh. 2009. Echinodermata (Asteroidea, Echinoidea and Holothuroidea) of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 209–220.
[CH97] Castro, P., & M. E. Huber. 1997. Marine Biology 2nd ed. WCB McGraw-Hill: Boston.
[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.
[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.
[SM15] Sampey, A., & L. M. Marsh. 2015. Kimberley marine biota. Historical data: echinoderms. Records of the Western Australian Museum Supplement 84: 207–246.
[SG93] Simms, M. J., A. S. Gale, P. Gilliland, E. P. F. Rose & G. D. Sevastopulo. 1993. Echinodermata. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 491–528. Chapman & Hall: London.
[SW66] Spencer, W. K., & C. W. Wright. 1966. Asterozoans. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt U. Echinodermata 3 vol. 1 pp. U4–U107. The Geological Society of America, Inc., and The University of Kansas Press.
[TS16] Thuy, B., & S. Stöhr. 2016. A new morphological phylogeny of the Ophiuroidea (Echinodermata) accords with molecular evidence and renders microfossils accessible for cladistics. PLoS One 11 (5): e0156140.
[W93] Wilson, B. 1993. Australian Marine Shells vol. 1. Prosobranch Gastropods. Part One. Odyssey Publishing: Kallaroo (Western Australia).