Belongs within: Caridea.
The Pandalidae are a cosmopolitan family of shrimp found primarily in colder waters (including deep waters in tropical regions).
Characters (from Chace 1985): Rostrum well-developed, laterally compressed, variable; antennular flagella simple, without accessory branches; mandible with palp and with incisor and molar processes deeply separate; first maxilliped with flagellum on exopod; second maxilliped normal, two distal segments not arising side by side from penultimate segment; first pereopod simple or microscopically chelate; second pereopod chelate, carpus subdivided into two or more articles; first male pleopod with endopod normal, not unusually enlarged or elaborately convoluted.
<==Pandalidae |--Chlorocurtis Kemp 1925C85 |--AustropandalusC85 |--Procletes levicarina (Bate 1888)MG-H11 |--Notopandalus Yaldwyn 1960B04 |--Dorodotes Bate 1888 [Dorodoteidae]C85 | `--*D. reflexus Bate 1888C85 |--Heterocarpoides De Man 1917 [Heterocarpoididae]C85 | `--*H. levicarina (Bate 1888) (see below for synonymy)C85 |--Stylopandalus Coutière 1905C85 | `--*S. richardi (Coutière 1905) (see below for synonymy)C85 |--Dichelopandalus Caullery 1896B04 | |--D. bonnieriCB93 | `--D. leptocerusK-M02 |--PandalinaC85 | |--P. brevirostris (Rathke 1843)KK03 | `--P. profunda Holthuis 1946KK03 |--Chlorotocella Balss 1914C85 | |--*C. gracilis Balss 1914C85 | `--C. spinicaudus (Milne Edwards 1837)MG-H11 |--Chlorotocus Milne Edwards 1882C85 | |--C. crassicornis (Costa 1871) (see below for synonymy)C85 | `--C. novaezeelandiae (Borradaile 1916)C85 |--Pandalus Leach 1814B04 | |--P. borealisMF02 | |--P. brevis Rathbun 1906C85 | |--P. jordaniB78 | |--P. montaguiAMI02 | |--P. punctulatus Risso 1826R26 | `--P. pelagicus Risso 1826R26 |--Heterocarpus Milne Edwards 1881 [Heterocarpidae]C85 | |--*H. ensifer Milne Edwards 1881C85 | | |--H. e. ensiferC85 | | `--H. e. parvispina De Man 1917C85 | |--H. affinis Faxon 1893C85 | |--H. alexandri Milne Edwards 1883C85 | |--H. dorsalis Bate 1888 [incl. H. affinis Borradaile 1915 non Faxon 1893, H. alphonsi Bate 1888]C85 | |--H. gibbosus Bate 1888C85 | |--H. grimaldii Milne Edwards & Bouvier 1900C85 | |--H. hayashii Crosnier 1988MG-H11 | |--H. hostilis Faxon 1893C85 | |--H. laevigatus Bate 1888C85 | |--H. laevis Milne Edwards 1883C85 | |--H. lepidus De Man 1917C85 | |--H. longirostris MacGilchrist 1905C85 | |--H. oryx Milne Edwards 1881C85 | |--H. reedi Bahamonde 1955C85 | |--H. sibogae De Man 1917C85 | |--H. signatus Rathbun 1906C85 | |--H. tricarinatus Alcock & Anderson 1894C85 | |--H. unicarinatus Borradaile 1915C85 | |--H. vicarius Faxon 1893C85 | `--H. woodmasoni Alcock 1901C85 `--Plesionika Bate 1888 [incl. Nothocaris Bate 1888, Parapandalus Borradaile 1899; Plesionikidae]C85 |--P. ensis (Milne Edwards 1881) [=Acanthephyra ensis; incl. *P. uniproducta Bate 1888]C85 |--P. acanthonotus (Smith 1882) [=Pandalus acanthonotus]C85 |--P. acinacifer Chace 1985C85 |--P. adensameri (Balss 1914)C85 |--P. alcocki (Anderson 1896) [=Pandalus alcocki]C85 |--P. antigai Zariquiey 1955C85 |--P. assimilis De Man 1917C85 |--P. beebei Chace 1937C85 |--P. bifurca Alcock & Anderson 1894 [=Pandalus bifurca]C85 |--P. binoculus (Bate 1888) [=Nothocaris binoculus]C85 |--P. brevirostris Bate 1888C85 |--P. costelloi (Yaldwyn 1971)C85 |--P. edwardsii (Brandt 1851) (see below for synonymy)C85 |--‘Pandalus’ escatilis Stimpson 1860C85 |--P. exigua (Rathbun 1906)C85 |--P. filipes (Calman 1939)C85 |--P. fimbriata Chace 1985C85 |--P. gigliolii (Senna 1903)BRP03 |--P. gracilis (Zarenkov 1971)C85 |--P. grandis Doflein 1902 [=Pl. spinipes var. grandis]C85 |--P. heterocarpus (Costa 1871)C85 |--P. holthuisi Crosnier & Forest 1968C85 |--P. hypanodon Doflein 1902C85 |--P. indica De Man 1917 [=P. longipes var. indica]C85 |--P. intermedia Chace 1985C85 |--P. izumiae Omori 1971C85 |--P. kensleyi Chace 1985C85 |--P. longipesC85 |--P. lophotes Chace 1985C85 |--P. martia (Milne Edwards 1883)C85 | |--P. m. martiaC85 | `--P. m. orientalis Chace 1985C85 |--P. mexicana Chace 1937C85 |--P. minor Calman 1939C85 |--P. multispinosus (Zarenkov 1971)C85 |--P. narval (Fabricius 1787) (see below for synonymy)C85 |--P. ocellus (Bate 1888) [=Nothocaris ocellus; incl. Pandalus sindoi Rathbun 1906, Plesionika sindoi]C85 |--P. orientalis Chace 1985MG-H11 |--P. ortmanni Doflein 1902C85 |--P. pacifica Edmondson 1952C85 |--P. parvimartia Chace 1985C85 |--P. persica (Kemp 1925)C85 |--P. philippinensis Chace 1985C85 |--P. pumila Chace 1985C85 |--P. quasigrandis Chace 1985C85 |--P. reflexa Chace 1985C85 |--P. rostricrescentis (Bate 1888) [=*Nothocaris rostricrescentis]C85 |--P. semilaevis Bate 1888 [=P. martia var. semilaevis]C85 |--P. serratifrons (Borradaile 1899) [=Pandalus (*Parapandalus) serratifrons]C85 |--P. simulatrix Chace 1985C85 |--P. spinensis Chace 1985C85 |--P. spinidorsalis (Rathbun 1906)C85 |--P. spinipes Bate 1888 [=Parapandalus spinipes]C85 |--P. tenuipes (Smith 1881) [=Pandalus tenuipes]C85 |--P. ‘tenuipes’ (Borradaile 1899) non Pandalus tenuipes Smith 1881C85 |--P. trispinus Squires & Barragan 1976C85 `--P. unidens Bate 1888C85
Chlorotocus crassicornis (Costa 1871) [=Pandalus crassicornis; incl. C. gracilipes var. andamanensis Alcock & Anderson 1899, *C. gracilipes Milne Edwards 1882, C. incertus Bate 1888 (n. d.)]C85
*Heterocarpoides levicarina (Bate 1888) [=Dorodotes levicarina, Heterocarpus (Heterocarpoides) levicarina; incl. Heterocarpus (Heterocarpoides) glabrus Zarenkov 1971]C85
Plesionika edwardsii (Brandt 1851) [=Pandalus (Pontophilus) edwardsii; incl. Pandalus (Parapandalus) longirostris Borradaile 1899]C85
Plesionika narval (Fabricius 1787) [=Astacus narval, Pandalus narval, Parapandalus narval; incl. Pl. pristis (Risso 1816)]C85
*Stylopandalus richardi (Coutière 1905) [=Pandalus richardi; incl. Pan. (Plesionika) gracilis Borradaile 1915, Parapandalus gracilis, Par. zurstrasseni Balss 1914]C85
*Type species of generic name indicated
References
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[BRP03] Bottari, T., P. Rinelli & A. Panebianco. 2003. Trawling lesions in some decapod crustaceans: considerations with regard to government inspections. Crustaceana 76 (8): 927–933.
[B04] Boyko, C. B. 2004. The Bopyridae (Crustacea, Isopoda) parasites of the Stylodactylidae (Crustacea, Decapoda, Caridea). Zoosystema 26 (2): 199–210.
[B78] Brusca, R. C. 1978. Studies on the cymothoid fish symbionts of the eastern Pacific (Crustacea: Isopoda: Cymothoidae) II. Systematics and biology of Lironeca vulgaris Stimpson 1857. Occasional Papers of the Allan Hancock Foundation New Series 2: 1–19.
[CB93] Cash, C. E., & R. T. Bauer. 1993. Adaptations of the branchial ectoparasite Probopyrus pandalicola (Isopoda: Bopyridae) for survival and reproduction related to ecdysis of the host, Palaemonetes pugio (Caridea: Palaemonidae). Journal of Crustacean Biology 13 (1): 111–124.
[C85] Chace, F. A., Jr. 1985. The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine expedition, 1907–1910, part 3: Families Thalassocarididae and Pandalidae. Smithsonian Contributions to Zoology 411: 1–143.
[K-M02] Klein-MacPhee, G. 2002. Silver hakes. Family Merlucciidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 217–222. Smithsonian Institute Press: Washington.
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[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.
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