Belongs within: Brachystigmata.
Contains: Libellulidae, Hemicorduliidae, Synthemistidae.
The Paneurypalpidomorpha are a clade of dragonflies in which the wings have the anterior margin of the hypertriangle distinctly convex, and the primary antenodals close together (Bechly & Ueda 2002).
Synapomorphies (from Bechly & Ueda 2002) Anterior margin of hypertriangle distinctly convex, especially in the hindwings, because of a basally arched MA; primary antenodal Ax1 in fore wing shifted basal of level of distal angle of discoidal triangle; Ax1 and Ax2 in both pairs of wings relatively close together with not more than one secondary antenodal crossvein between them; lestine oblique vein only 1–2 cells distad of subnodus in both pairs of wings; forewing discoidal triangle more or less transverse instead of elongate; discoidal triangles unicellular in both pairs of wings (occasionally reversed in hind wing).
<==PaneurypalpidomorphaBU02
|--Juracordulia Bechly 1998 [Juracorduliidae]BU02
| `--J. schiemenzi Bechly 1998BU02
`--EurypalpidomorphaBU02
|--ValdicordulioideaBU02
| |--ValdicorduliidaeBU02
| `--AraripephlebiidaeBU02
`--EurypalpidiformiaBU02
|--EocorduliaR02 [EocorduliidaeBU02]
| `--E. cretacea Pritykina 1986RJ93
`--Libelluloidea [Paneurypalpida]BU02
| i. s.: PalaeolibellulaS02
|--LibellulidaeTH06
|--HemicorduliidaeWO91
|--SynthemistidaeWO91
|--OxygastridaeTH06
| |--Hesperocordulia berthoudi Tillyard 1911WO91, TH06
| `--OxygastraTH06
|--Pseudocordulia Tillyard 1909T09 [PseudocorduliidaeTH06]
| |--*P. circularis Tillyard 1909T09
| `--P. elliptica Tillyard 1913TH06
|--Gomphomacromiidae [Gomphomacromiinae]WO91
| |--Gomphomacromia Brauer 1864FB03
| | |--*G. paradoxaT09
| | `--G. volxemiT09
| `--ArchaeophyaTH06
| |--A. adamsi Fraser 1959TH06
| `--A. magnifica Theischinger & Watson 1978TH06
|--Cordulephyidae [Cordulephyinae]FB03
| |--NeocorduliaTH06
| `--CordulephyaTH06
| |--*C. pygmaea Selys 1870T09, TH06
| |--C. bidens Sjöstedt 1917TH06
| |--C. divergens Tillyard 1917TH06
| `--C. montana Tillyard 1911TH06
|--Urothemistidae [Macrodiplactidae, Urothemistinae]TH06
| |--Macrodiplax cora (Kaup 1867)TH06
| |--UrothemisTH06
| | |--U. aliena Selys 1878TH06
| | `--U. sanguinea (Müll. 1767)SL89, L49 [=Libellula sanguineaSL89; incl. L. roeseliiL49, L. signataSL89]
| `--AethriamantaTH06
| |--A. circumsignata Selys 1897TH06
| `--A. nymphaeae Lieftinck 1949TH06
|--Macromiidae [Macromiinae]GE05
| |--Pseudomacromia [Pseudomacromiidae]GE05
| | `--P. sensibilis Carle & Wighton 1990RJ93
| `--MacromiaTH06
| |--M. cinctaSL89
| |--M. clio Ris 1916I92
| |--M. kubokaiya Asahina 1964I92
| |--M. magnificaA99
| |--M. splendensT09
| |--M. tillyardi Martin 1906TH06
| |--M. urania Ris 1916I92
| |--M. viridescens Tillyard 1911TH06
| `--M. westwoodiiSL89
|--Corduliidae [Cordulinae]WO91
| |--Pentathemis membranulata Karsch 1890TH06
| |--Metaphya tillyardi Ris 1913TH06
| |--DorocorduliaA07
| |--EpithecaP74
| |--HelocorduliaP74
| |--NeurocorduliaP74
| |--SomatochloraA07
| |--Cordulia aeneaK01
| |--DidymopsA07
| |--EpicorduliaA07
| |--TetragoneuriaA07
| |--*Neophya rutherfordiT09
| `--SyncorduliaT09
`--AustrocorduliidaeTH06
|--Apocordulia macrops Watson 1980TH06
|--Austrophya Tillyard 1909TH06, T09
| `--*A. mystica Tillyard 1909T09
|--LathrocorduliaTH06
| |--L. garrisoni Theischinger & Watson 1991TH06
| `--L. metallica Tillyard 1911TH06
|--MicromidiaTH06
| |--M. atrifrons (McLachlan 1883)TH06
| |--M. convergens Theischinger & Watson 1978TH06
| `--M. rodericki Fraser 1959TH06
`--Austrocordulia Tillyard 1908T09
|--*A. refracta Tillyard 1909T09
| |--A. r. refractaTH06
| `--A. r. jurzitzaiTH06
|--A. leonardi Theischinger 1973TH06
`--A. territoria Theischinger & Watson 1978TH06
*Type species of generic name indicated
References
[A07] Anderson, T. M. 2007. An assessment of water mite parasitism of dragonflies based on museum collections. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 61–69. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.
[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (translated: 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).
[BU02] Bechly, G., & K. Ueda. 2002. The first fossil record and first New World record for the dragonfly clade Chlorogomphida (Insecta: Odonata: Anisoptera: Araripechlorogomphidae n. fam.) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 328: 1–11.
[FB03] Fleck, G., G. Bechly, X. Martínez-Delclòs, E. Jarzembowski, R. Coram & A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France 39 (1): 55–93.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.
[K01] Klapálek, F. 1901. Neuropteroidák [Neuropteroiden]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 203–221. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.
[P74] Paulson, D. R. 1974. Reproductive isolation in damselflies. Systematic Zoology 23 (1): 40–49.
[R02] Rasnitsyn, A. P. 2002. Special features of the study of fossil insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 8–12. Kluwer Academic Publishers: Dordrecht.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
[SL89] Selys Longchamps, E. de. 1889. Odonates de Sumatra comprenant les espèces recueillies à Pulo Nias par M. le D.r E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 444–484.
[S02] Sinitshenkova, N. D. 2002. Ecological history of the aquatic insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 388–426. Kluwer Academic Publishers: Dordrecht.
[TH06] Theischinger, G., & J. Hawking. 2006. The Complete Field Guide to Dragonflies of Australia. CSIRO Publishing: Collingwood (Victoria).
[T09] Tillyard, R. J. 1909. On some remarkable Australian Corduliinae, with descriptions of new species. Proceedings of the Linnean Society of New South Wales 33: 737–751, pls 21–22.
[WO91] Watson, J. A. L., & A. F. O’Farrell. 1991. Odonata (dragonflies and damselflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 294–310. Melbourne University Press: Carlton (Victoria).