Belongs within: Peripsocoidea.
The Peripsocidae are a group of barklice that are most common on bark or dead foliage, more rarely on low vegetation (New & Lienhard 2007). Genera of Peripsocidae are very similar in appearance and can often be distinguished only by genitalic features.
Characters (from New & Lienhard 2007): Fore wing length ca 1.8–2.4 mm. Lacinial apex slender, bicuspid, outer cusp usually broader than inner. Vertex smooth or with few small setae. Tarsi two-segmented; claws with subapical tooth, pulvillus slender. Fore wing lacking areola postica; pterostigma of normal form, more or less expanded in central region. Wing setation usually reduced or absent, occasionally more conspicuous; hind wing glabrous, veins Rs and M fused for a length. Male with hypandrium simple; phallosome a closed frame, with pair of poriferous parameres; aedeagal arch usually with distinct apical constriction or process; endophallus with symmetrical sclerites. Female subgenital plate often with distinct, commonly bilobed, apical region; gonapophyses complete; dorsal valve large, usually setose at apex; external valve large, setose.
<==Peripsocidae [Peripsocinae] |--Orbiperipsocus Li 2002 [=Orbiperipaocus]L03 | `--*O. fractiflexus Li 2002L03 |--Campanulatus Li 2002 [=Canpanulatus]L03 | |--*C. jaculatorius Li 2002 [=C. jaculotoria, C. jaculotorius]L03 | |--C. lagenarius Li 2002L02 | `--C. pictus (Thornton 1962) [=Peripsocus pictus]L02 |--Bicuspidatus Li 1993L02 | |--*B. pulchipunctatus Li 1993L02 | |--B. guangdongensis Li 1993L02 | `--B. sigillatus Li 1993L02 |--Properipsocus Li 1995L02 | |--*P. gracilis Li 1995L02 | |--P. bicornis (Thornton 1962) [=Peripsocus bicornis, Periterminalis bicornis]L02 | `--P. quadartus Li 2002 [=P. qradartus]L03 |--Kaestneriella Roesler 1943M93 | |--*K. pilosaM93 | |--K. fumosa (Banks 1903) [=Peripsocus fumosus]M93 | |--K. guatemalensisY02 | `--K. tenebrosa Mockford & Sullivan 1990M93 |--Pericupsocus Li 2002L02 | |--*P. cuspidatus Li 2002L02 | |--P. alboguttatus (Dalman 1823)L02 | |--P. crenulatus (Thornton & Wong 1968)L02 | `--P. digitalis Li 2002L02 |--Cycloperipsocus Li 1993SN08 [=Cycloperopsocus (l. c.)L03] | |--*C. pangi Li 1993SN08 | |--C. diplocyclus Li 2002L02 | |--C. edwardsi (New 1973) [=Peripsocus edwardsi]SN08 | `--C. plurimaculatus (Li 1992) [=Peripsocus plurimaculatus]L02 |--Periterminalis Li 1997L02 [=Periteriminalis (l. c.)L03] | |--*P. crytomeriae Li 1997L02 | |--P. hiatus (Thornton & Wong 1968) [=Peripsocus hiatus]NL07 | |--P. lata Li 2002 [=P. latus]L03 | |--P. longicuspis Li 2002L02 | `--P. shanxiensis Li 2002L02 |--Turriperipsocus Li 1997L02 | |--*T. cunninghamiae Li 1997L02, L03, L02 [=Peripsocus cunninghamiusL02, T. cunninghamius (l. c.)L03] | |--T. chiangensis (New 1973) [=Peripsocus chiangensis]NL07 | |--T. cupressisugus Li 2002L02 | |--T. decimidentatus Li 1997L02 | |--T. hypsodontus Li 2002L02 | `--T. leshanensis Li 2002L02 |--Coniperipsocus Li 1997L02 [=Comiperipsocus (l. c.)L03] | |--*C. yuae Li 1997L02 | |--C. jindingensis Li 2002L03 | |--C. melanolomus Li 2002L02 | |--C. minutissimus Li 2002L02 | |--C. proceridentalis Li 2002L02 | |--C. quadrangulus (Li 1992) [=Diplopsocus quadrangulus]L03 | `--C. quadrifascius (Li & Mockford 1993) [=Diplopsocus quadrifascius]L02 |--Diplopsocus Li & Mockford 1993L03 | |--*D. cupressicolus Li & Mockford 1993L02 [=D. cupressicola (l. c.)L03] | |--D. albostigmus Li & Mockford 1993L02 | |--D. anoplus (Thornton & Wong 1968)NL07 | |--D. chebalingicus Li 1993L02 | |--D. circularis (New & Thornton 1976)NL07 | |--D. fasciatus (Thornton 1959)NL07 | |--D. fujianicus Li & Mockford 1993L02 | |--D. hainanensis Li & Mockford 1993L02 | |--D. huananensis Li 2002L02 | |--D. irregularis Li 1995L02 | |--D. jiangxiensis Li & Mockford 1993L02 | |--D. liupanshanensis Li 2002 [=D. liupenshanensis]L03 | |--D. lophacanthus Li & Mockford 1993L02 | |--D. magniocellatus Li 1993L03 | |--D. melanostictus Li & Mockford 1993L02 | |--D. ningmingicus Li & Mockford 1993L02 | |--D. oblongus Li & Mockford 1993L02 | |--D. palmiformis Li & Mockford 1993L02 | |--D. parabolicus Li & Mockford 1993L02 | |--D. parviocellatus Li 1993L02 | |--D. phaeophlebicus Li 2002 [=D. phaeophlebius]L03 | |--D. phagococcus Li & Mockford 1993L02 | |--D. philopinnus Li & Mockford 1993L02 | |--D. porosus Li & Mockford 1993L02 | |--D. resupinatus Li & Mockford 1993L02 | |--D. rhombeus Li & Mockford 1993L02 | |--D. scrobiculatus Li 2002L02 | |--D. selene (Thornton & Wong 1968)NL07 | |--D. shanxiensis Li & Mockford 1993L02 | |--D. sichuanicus Li 1997L02 | |--D. sylvaticus Li 2002L02 | |--D. tibeticus Li & Mockford 1993L02 | |--D. xilingxiaensis Li 1997L02 | `--D. yangi Li & Mockford 1993L02 `--Peripsocus Hagen 1866SN08 [incl. Peripsocopsis Tillyard 1923L98] | i. s.: P. africanus Enderlein 1902E07 | P. ammonus Li 2002L02 | P. annectens Li 2002L02 | P. anura Vaughan, Thornton & New 1991NL07 | P. apicatus Vaugan, Thornton & New 1989NL07 [=Properipsocus apicatusL02] | P. apiculatus Li 2002L02 | P. attenuatus Li 2002L02 | P. badimaculatus Li 1993L02 | P. baishanzuicus Li 1995L02 | P. baiyunshanicus Li 2002L02 | P. barunus Vaughan, Thornton & New 1991NL07 | P. beijingensis Li 2002L02 | P. biacanthus Li & Yang 1988L02 | P. bifasciarius Li 2002L02 | P. bifasciatusSN08 | P. bifidus Thornton 1984NL07 | P. bivari Baz 1988L98 | P. brachyura Thornton & Wong 1968NL07 | P. brinchangensis New & Lee 1991NL07 | P. bucephalus Li 1992L02 | P. bucerus Li 1992L02 | P. bulbus Li 2002L02 | P. camerunusB55 | P. cassideus Li 1999L02 | P. caudatus Li 1997L02 | P. changbaishanicus Li 2002L02 | P. circinus Thornton & Wong 1968NL07 | P. cochleus Schmidt & New 2008SN08 | P. conoidalis Li 2002L02 | P. consobrinus Pearman 1951L98 | P. constrictus Thornton & Wong 1968NL07 | P. corollaris Li 2002L02 | P. crassicosta New & Lee 1991NL07 | P. crenulatus Thornton & Wong 1968NL07 | P. cylindratus Li 2002L02 | P. decurvatus Li 1995L02 | P. denticulatus Thornton & Wong 1968NL07 | P. disdentus Li 1997L03, L02 [=P. distentus (l. c.)L03] | P. dongbeiensis Li 2002L02 | P. duodecimidentus Li 2002L02 | P. elongatus New & Thornton 1976NL07 | P. equispineus Li 2002 [=P. eguispineus]L03 | P. exilis Li 1997L02 | P. falsipictus Li 2002L02 | P. ferrugineus Thornton & Wong 1968NL07 | P. fici Smithers 1984T13 | P. forcipatus Li 1993L02 | P. forficatus Li 2002L02 | P. fornicalis Li 1999L02 | P. frimensis New & Lee 1991NL07 | P. fulvescens Navás 1920NL07 | P. furcellatus Li 2002L02 | P. grandispineus Li 2002L02 | P. guandishanicus Li 2002L02 | P. hainanensis Li 2002L02 | P. haplacanthus Li 1993L02 | P. hedinianus Enderlein 1936L02 | P. hickmaniSN08 | P. hongkongensis Thornton & Wong 1968NL07 | P. huashanensis Li 2002L02 | P. ignis Okamoto 1910NL07 | P. jianfenglingicus Li 2002L02 | P. jiangxiensis Li 2002L02 | P. jilinicus Li 2002L02 | P. jinggangshanicus Li 2002L02 | P. jinshaanensis Li 2002L02 | P. jinxiuensis Li 2002L02 | P. kunmingiensis Li 2002L02 | P. laoshanicus Li 2002L02 | P. laricis Li 2002L02 | P. latispineus Li 2002L02 | P. leleupi Badonnel 1976L98 | P. leptorrhizus Li 1997L02 | P. lifashengi Lienhard 2003 [=P. orientalis Li 2002 non Garcia Aldrete 1999]L03 | P. limi New & Thornton 1976NL07 | P. longifurcus Li 2002L02 | P. louguantaiensis Li 2002L02 | P. lunaris Li 1993L02 | P. luotongshanicus Li 2002L02 | P. machadoi Badonnel 1955B55 | P. macrosiphus Li 1993L02 | P. magnimammus Li 2002L02 | P. maoricus (Tillyard 1923)SN08 (see below for synonymy) | P. medifasciarius Li 2002 [=P. medifaciarius]L03 | P. medimacularis Li 1995L02 | P. medispineus Li 2002L02 | P. megalophus Li 1995L02 | P. melaleucae New 1971SN08 | P. meridionalis Li 1999L02 | P. microcheilus Li 1993 [=P. microcheilius]L03 | P. milleri (Tillyard 1923) [=Peripsocopsis milleri; incl. Peripsocus eucalypti Edwards 1950]SN08 | P. mingshanicus Li 1997L02 | P. mirabilis Li 1995L02 | P. morulops (Tillyard 1923) [=Peripsocopsis morulops]SN08 | P. mutilatus Badonnel 1986B86 | P. nanjingensis Li 2002L02 | P. nanus Navás 1922NL07 | P. nasutusB86 | P. nitens Thornton & Wong 1968NL07, L98 | P. octoidentus Li 2002L02 | P. oculatus Enderlein 1926NL07 | P. ‘oculatus’ Badonnel 1955 non Enderlein 1926 [=P. ocuatus]B55 | P. oculimacularis Li 2002L02 | P. odontopetalus Li 2002 [=P. codntopetalus]L03 | P. oligodontus Li 2002L02 | P. optimalis Li 1997L02 | P. orbiculatus Li 1993L02 | P. orebius Li 2002L02 | P. orientalis Garcia Aldrete 1999NL07 | P. oxydontus Li 2002L02 | P. pamae Schmidt & New 2008SN08 | P. papillatus Vaughan, Thornton & New 1989NL07 | P. parareicherti Vaughan, Thornton & New 1989NL07 | P. paraspinosus New & Lee 1991NL07 | P. parvulus Kolbe 1880 [=P. alboguttatus var. parvulus; incl. P. parvulus f. longipennis Roesler 1939]L98 | P. parvus Li 1995L02 | P. phaeochilus Li 2002L02 | P. plagiotropus Li 2002L02 | P. platyopterus Li 1997L02 | P. plstypus Li 1995L03, L02 [=L. platylpus (l. c.)L03, L. platypus (l. c.)L03] | P. polygonalis Li 1995L02 | P. polyoacanthus Li 1993L02 | P. pseudoquercicolus Thornton 1962L02 | P. qingchengshanicus Li 2002L02 | P. qingdaoensis Li 1993L02 | P. quadratidentalis Li 2002L02 | P. quadratiprocessus Li 1993L02 | P. quadripunctatus Badonnel 1955L02 [incl. P. quadripunctatus f. brachypterus Badonnel 1955B55] | P. quattuordecimus Li 1995L02 | P. quercicola Enderlein 1906NL07 | P. reduncus Li 2002L02 | P. reflexibilis Li 2002L02 | P. reicherti Enderlein 1903NL07 | P. rhombicus Li 2002L02 | P. rhomboacanthus Li 2002 [=P. rhombocanthus]L03 | P. scalpratus Li 2002L02 | P. scapiformis Li 2002L02 | P. sedcimidentalis Li 1997 [=P. sedecimidentalis]L03 | P. sexidentus Li 2002L02 | P. shilinensis Li 2002L02 | P. siculiformis Li 2002L02 | P. ‘similis’ Enderlein 1903 non P. phaeopterus var. similis Kolbe 1880NL07 | P. ‘similis’ Badonnel 1955 nec Kolbe 1880 nec Enderlein 1903B55 | P. singularis Banks 1937L02 | P. spinosus Thornton 1959L02 | P. stenopterus Thornton & Wong 1964L02 | P. stigmatus Thornton & Wong 1968NL07 | P. stigmostigmus Li 2002L02 | P. stipiatus Li 1993L02 | P. suffitus Enderlein 1903NL07 | P. suoxiyuicus Li 2002L02 | P. taipingensis New & Lee 1991NL07 | P. terricolis Badonnel 1986L02 | P. tillyardi New 1973SN08 | P. tinctus Vaughan, Thornton & New 1989NL07 | P. transivenus Li 2002L02 | P. tredecimus Li 1997L02 | P. trigonoispineus Li 2002L02 | P. umbrosus Navás 1911 (n. d.)L98 | P. undecimidentus Li 2002L02 | P. undulatus Li 2002L02 | P. uniformis New & Thornton 1976NL07 | P. valvulus Thornton & Wong 1968NL07 | P. variatus Soehardjan & Hamann 1959NL07 [=Cycloperipsocus variatusL02] | P. varidentatus Li 2002 [=P. septemidentus Li 2001 (n. n.), P. varidentus]L03 | P. vescus Li 1997L02 | P. viriosus Li 1995L02 | P. wuhoi Li 2002L02 | P. wuyishanicus Li 1999L02 | P. xanthochilus Li 2002L02 | P. xihuensis Li 2002L02 | P. yongi New & Lee 1991NL07 | P. yuleki Galil in Galil & Halperin 1983L98 | P. zhangispineus Li 2002L02 | P. zhangliani Li 2002 [=P. zhangliangi, P. zhengliangi]L03 | P. ziguiensis Li 1997L02 |--Group IIIM93 | |--P. stagnivagus Chapman 1930M93 | `--Subgroup BM93 | |--*P. phaeopterus (Stephens 1836)M93 (see below for synonymy) | `--P. didymus Roesler 1939M93, L98 (see below for synonymy) `--Group IM93 |--Subgroup AM93 | |--P. madidus (Hagen 1861) [=Psocus madidus; incl. Ps. permadidus Walsh 1863, Caecilius permadidus]M93 | |--P. nigrescensM93 | |--P. pauliani Badonnel 1949M93 | |--P. quadrifasciatus (Harris 1869)M93 (see below for synonymy) | `--P. reductus Badonnel 1943M93 `--Subgroup BM93 |--P. alachuae Mockford 1971M93 |--P. alboguttatus (Dalman 1823)M93 (see below for synonymy) |--P. maculosus Mockford 1971M93 |--P. madescens (Walsh 1863) [=Psocus madescens]M93 |--P. minimus Mockford 1971M93 |--P. potosi Mockford 1971M93 `--P. teutonicusM93
Peripsocus alboguttatus (Dalman 1823)M93 [=Psocus alboguttatusL98; incl. Ps. pupillatus Walker 1853L98, Peripsocus pupillatusM93, Ps. quadrimaculatus Stephens 1836 non Latreille 1794L98]
Peripsocus didymus Roesler 1939M93, L98 [incl. P. didymus silesiaca Obr 1948L98, P. didymus truncatus Badonnel 1943L98]
Peripsocus maoricus (Tillyard 1923)SN08 [=*Peripsocopsis maoricusNL07, SN08; incl. Peripsocus macropterus Edwards 1950SN08]
*Peripsocus phaeopterus (Stephens 1836)M93 [=Psocus phaeopterusSN08; incl. Pe. fortunatus Navás 1916L98, Ps. nigricornis Stephens 1836L98, Ps. obscurus Rambur 1842L98, Peripsocus phaeopterus var. similis Kolbe 1880G74]
Peripsocus quadrifasciatus (Harris 1869)M93 [=Psocus quadrifasciatusL98; incl. Pe. subpupillatus ab. quadriramosus Enderlein 1901L98, Ps. subfasciatus Rambur 1842 non Zetterstedt 1840L98, Pe. subfasciatusL98, Pe. subpupillatus McLachlan 1883L98]
*Type species of generic name indicated
References
[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.
[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.
[E07] Enderlein, G. 1907. Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostafrikas 1905–1906 vol. 15. Corrodentia pt 2. Copeognatha. Almqvist & Wiksells Boktryckeri-A. B.: Uppsala.
[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.
[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.
[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.
[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.
[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.
[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.
[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.
[T13] Taylor, C. K. 2013. Annotated bibliography for Barrow Island terrestrial invertebrates. Records of the Western Australian Museum Supplement 83: 135–144.
[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.