Permochoristidae

Belongs within: Panorpoidea.

The Permochoristidae are scorpionfly-like insects known from the Permian to the Jurassic (Novokshonov 2002). Wings have SC bearing two anterior branches, a long MA fork, and five-branched MP (Bashkuev 2011).

<==Permochoristidae [Caenoptilonidae, Eosetidae, Idelopanorpidae, Petromantidae, Protochoristidae, Tychtopsychidae]
    |--PermochoristinaeB11
    |    |--Permochorista sinuata Tillyard 1922GE05, F71
    |    |--ProchoristellaB11
    |    |    |--P. anagaura Riek 1953F71
    |    |    |--P. belli (Tillyard 1926) [=Permochorista belli]F71
    |    |    |--P. concinna Riek 1953F71
    |    |    |--P. exilis Riek 1953F71
    |    |    |--P. leongatha Jell & Duncan 1986RJ93
    |    |    |--P. megaloprepia Riek 1953F71
    |    |    `--P. pusilla Riek 1953F71
    |    `--PermecaB11
    `--PseudonannochoristinaeRL09
         |--Neochoristella Riek 1953B11
         |    `--N. aptata Riek 1953F71
         |--Nannochoristella Riek 1953B11
         |    `--N. reducta Riek 1953F71
         |--PseudonannochoristaB11
         `--MiomecaB11

Permochoristidae incertae sedis:
  PetromantisGE05
  MesotanyderusR70
  LiassochoristaRL09
    |--L. anglicanaRJ93
    `--L. asiaticaN02
  MesagetaB11
  Mesochoristini [Mesochoristidae]B11
    |--MesochoristaKIR02
    |    |--M. australica (Tillyard 1917) (see below for synonymy)F71
    |    |--M. dubia Riek 1953F71
    |    |--M. javorskyiRJ93
    |    |--M. jucunda (Tillyard 1926) [=Permochorista jucunda]F71
    |    |--M. phipa Riek 1953F71
    |    |--M. proavita Tillyard 1916F71
    |    `--M. proavitella [incl. Eoses triassica]KIR02
    `--Baianochorista mongolica (Novokshonov 1977)B11

Mesochorista australica (Tillyard 1917) [=Permochorista australica; incl. P. affinis Tillyard 1922, P. angustipennis Tillyard 1926, P. collinsi Tillyard 1926, P. inaequalis Tillyard 1926, P. mitchelli Tillyard 1917, P. pincombei Tillyard 1926]^F71

*Type species of generic name indicated

References

[B11] Bashkuev, A. S. 2011. Nedubroviidae, a new family of Mecoptera: the first Paleozoic long-proboscid scorpionflies. Zootaxa 2895: 47–57.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[KIR02] Kozlov, M. V., V. D. Ivanov & A. P. Rasnitsyn. 2002. Order Lepidoptera Linné, 1758. The butterflies and moths (=Papilionida Laicharting, 1781). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 220–227. Kluwer Academic Publishers: Dordrecht.

[N02] Novokshonov, V. G. 2002. Order Panorpida Latreille, 1802. The scorpionflies (=Mecaptera Packard, 1886, =Mecoptera Comstock et Comstock, 1895, +Neomecoptera Hinton, 1958, +Paratrichoptera Tillyard, 1919, +Paramecoptera Tillyard, 1919). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 194–199. Kluwer Academic Publishers: Dordrecht.

[RL09] Ren, D., C. C. Labandeira, J. A. Santiago-Blay, A. Rasnitsyn, C. Shih, A. Bashkuev, M. A. V. Logan, C. L. Hotton & D. Dilcher. 2009. A probable pollination mode before angiosperms: Eurasian, long-proboscid scorpionflies. Science 326: 840–847.

[R70] Riek, E. F. 1970. Fossil history. In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 168–186. Melbourne University Press.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.