Belongs within: Hypnales.
Contains: Ectropothecium, Isopterygium, Vesicularia.
The Plagiotheciaceae are a family of mosses found worldwide, species of which grow in loose to dense mats or tufts. Though currently more diverse in the northern temperate zone, phylogenetic analysis indicates that the family may have originated in the Southern Hemisphere (Pedersen & Hedenäs 2002). Plagiothecium species are irregularly branching mosses with nerveless or two-nerved leaves and slender, oblong-cylindric capsules (Dixon 1924).
Synapomorphies (from Pedersen & Hedenäs 2002): Pseudoparaphyllia absent; rhizoids absent from stem and lowermost abaxial costa; axillary rhizoids purplish and granular-papillose; exostomes with whitish yellow basal part.
<==Plagiotheciaceae [Stereodontaceae] |--StereodonteaeBW15 | |--EctropotheciumBW15 | `--StereodonD03 | |--S. lechleriD03 | |--S. nitidusD03 | `--S. ptychocarpon [=Ptychomnion ptychocarpon, S. ptychomnium (l. c.)]SK02 `--PlagiothecieaeBW15 |--IsopterygiumBW15 |--VesiculariaBW15 |--Plagiotheciopsis oblonga [=Ectropothecium oblongum Brotherus 1895, Vesicularia oblonga Brotherus 1908]BW15 `--Plagiothecium Schimp. in Bruch, Schimp. & Gümbel 1851S04 [incl. Buckiella Ireland 2001OZB-O03] | i. s.: P. draytoniiOZB-O03 | P. fuegianumD03 | P. lamprostachys (Hampe) Jaeger 1878 (see below for synonymy)SK02 | P. leptoplumosum Dusén 1903D03 | P. lucidulumD03 | P. lucidum (Hooker & Wilson) Paris 1897SK02 | P. magellanicumD03 | P. neckeroideumN02 | P. piliferum (Schwartz) Schimp. in Bruch et al. 1851S04 (see below for synonymy) | P. subulatumB-OO10 | P. undulatum (Hedw.) Schimp. in Bruch et al. 1851S04 (see below for synonymy) |--P. (sect. Plagiothecium) denticulatum (Hedw.) Schimp. in Bruch, Schimp. & Gümbel 1851OZB-O03, S04 (see below for synonymy) | |--P. d. var. denticulatumS04 | |--P. d. var. majus [incl. P. rufo-virescens]D24 | |--P. d. var. obtusifolium (Turner) Moore 1873S04 | `--P. d. var. undulatum Ruthe ex Geheeb 1877S04 (see below for synonymy) |--P. sect. Leptophyllum Jedl. 1948OZB-O03 | |--P. curvifolium Schlieph. ex Limpr. 1897OZB-O03, S04 (see below for synonymy) | `--P. laetum Schimp. in Bruch, Schimp. & Gümbel 1851OZB-O03, S04 (see below for synonymy) |--P. sect. Orthophyllum Jedl. 1948OZB-O03 | |--P. cavifolium (Brid.) Iwats. 1970OZB-O03, S04 (see below for synonymy) | |--P. nemorale (Mitt.) Jaeger 1878OZB-O03, S04 (see below for synonymy) | | |--P. n. var. nemoraleD24 | | `--P. ‘silvaticum’ var. sullivantiaeD24 | `--P. succulentum (Wilson) Lindb. 1865OZB-O03, S04 (see below for synonymy) |--P. sect. Philoscia (Berk.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 (see below for synonymy)OZB-O03 | `--P. latebricola Schimp. in Bruch, Schimp. & Gümbel 1851OZB-O03, S04 (see below for synonymy) | |--P. l. f. latebricolaD24 | `--P. l. f. gemmascensD24 `--P. sect. Rostriphyllum Jedl. 1948OZB-O03 `--P. platyphyllum Mönk. 1927OZB-O03, S04 (see below for synonymy) Plagiotheciaceae incertae sedis: Catagonium Müll.Hal. 1908SK02 `--C. nitens (Brid.) Cardot 1915 (see below for synonymy)SK02 Acrocladium Mitt. 1869SK02 |--A. auriculatum [incl. Hypnum auriculatum, Acroceratium auriculatum, Stereodon auriculatus]SK02 |--A. chlamydophyllum (Hooker & Wilson) Müll.Hal. & Broth. 1900 (see below for synonymy)SK02 |--A. morenoiD03 `--A. trichocladium Bosw. 1892SK02 Myurella Schimp. in Bruch, Schimp. & Gümbel 1853 non Hinds 1845 (ICZN)S04 |--M. careyanaD24 |--M. julacea (Schwägr.) Schimp. in Bruch et al. 1853S04 (see below for synonymy) | |--M. j. var. julaceaOZB-O03 | `--M. j. var. ciliata (Chal.) Ochyra & Bednarek-Ochyra 1991 [=M. apiculata var. ciliata]OZB-O03 `--M. tenerrima (Brid.) Lindb. 1879S04 [incl. M. apiculataS04, Isothecium apiculatumD24] Orthothecium Schimp. in Bruch, Schimp. & Gümbel 1851S04 |--O. chryseon (Schwägr.) Schimp. in Bruch, Schimp. & Gümbel 1851 (see below for synonymy)OZB-O03 |--O. intricatum Schimp. in Bruch, Schimp. & Gümbel 1851S04 (see below for synonymy) | |--O. i. var. intricatumD24 | `--O. i. var. binervulumD24 `--O. rufescens (Dicks. ex Brid.) Schimp. in Bruch, Schimp. & Gümbel 1851S04 (see below for synonymy) Pseudotaxiphyllum Iwats. 1987S04 `--*P. elegans (Brid.) Iwats. 1987OZB-O03, S04 (see below for synonymy) |--P. e. var. elegansD24 `--‘Plagiothecium’ e. var. collinumD24 Herzogiella Broth. 1925S04 [incl. Dolichotheca (nom. illeg.)OZB-O03, Sharpiella Iwatsuki 1965OZB-O03] |--H. cylindricarpaJ87 |--H. seligeri (Brid.) Iwats. 1970S04 (see below for synonymy) `--H. striatella (Brid.) Iwats. 1970S04 (see below for synonymy) Taxiphyllum Fleischer 1923S04, SK02 |--T. densifolium (Lindb. ex Broth.) Reimers 1940 (see below for synonymy)OZB-O03 |--T. taxirameum (Mitt.) Fleischer 1923 (see below for synonymy)SK02 `--T. wissgrillii (Garov.) Wijk & Margad. 1960S04 (see below for synonymy) Isopterygiopsis Iwats. 1970S04 |--*I. muelleriana (Limpr.) Iwats. 1970OZB-O03, S04 [=Isopterygium muellerianumS04, Plagiothecium muellerianumS04] `--I. pulchella (Hedw.) Iwats. 1987S04 (see below for synonymy) Nomen nudum: Plagiothecium amblyostomum Müll.Hal. ex Burges 1935SK02 Stereodon australis Hooker & Wilson ex Muell. 1880SK02 Stereodon cupressiformis f. robusta Rodway 1914SK02 Stereodon wattsii Broth. ex Watts 1899SK02
Acrocladium chlamydophyllum (Hooker & Wilson) Müll.Hal. & Broth. 1900 [=Hypnum chlamydophyllum, Coelidium chlamydophyllum, Lembophyllum chlamydophyllum]SK02
Catagonium nitens (Brid.) Cardot 1915 [incl. Phyllogonium elegans var. acuminatum (nom. inv.), Orthorrhynchium hampeanum, Cryptogonium hampeanum, Phyllogonium hampeanum, Orthorrhynchium hampei (n. n.), Hypnum politum, Acroceratium politum, Acrocladium politum, Catagonium politum]SK02
Herzogiella seligeri (Brid.) Iwats. 1970S04 [=Dolicotheca seligeriS04, Hypnum seligeriOZB-O03, Isopterygium seligeriS04, Leskia seligeriOZB-O03, Plagiothecium seligeriOZB-O03, Sharpiella seligeriS04; incl. Dolichotheca repensOZB-O03, Isopterygium repensD24, Plagiothecium repensOZB-O03, Dolichotheca silesiacaOZB-O03, Hypnum silesiacumOZB-O03, Isopterygium silesiacumOZB-O03, Plagiothecium silesiacumD24]
Herzogiella striatella (Brid.) Iwats. 1970S04 [=Campylium striatellumOZB-O03, Dolichotheca striatellaS04, Hypnum striatellumOZB-O03, Isopterygium striatellumOZB-O03, Leskia striatellaOZB-O03, Plagiothecium striatellumS04, Sharpiella striatellaS04; incl. Hypnum muehlenbeckiiD24, Plagiothecium muehlenbeckiiS04]
Isopterygiopsis pulchella (Hedw.) Iwats. 1987S04 [=Hypnum pulchellumD24, Isopterygium nitidum var. pulchellumOZB-O03, Isopterygium pulchellum (Hedw.) Jaeger 1878SK02, Leskea pulchellaOZB-O03, Plagiothecium nitidum var. pulchellumOZB-O03, Stereodon pulchellusOZB-O03; incl. Hypnum nitidulum (nom. illeg.)SK02, Isopterygium pulchellum var. nitidulumD24, S04, Leskea nitidulaOZB-O03, Plagiothecium pulchellum var. nitidulumD24, Isopterygium nitidumOZB-O03, P. nitidumOZB-O03, P. nitidum var. suberectumOZB-O03]
Myurella julacea (Schwägr.) Schimp. in Bruch et al. 1853S04 [=Hypnum julaceumD24, Leskea julaceaOZB-O03; incl. Myurella julacea var. scabrifoliaS04]
Orthothecium chryseon (Schwägr.) Schimp. in Bruch, Schimp. & Gümbel 1851 [=Holmgrenia chrysea, Hypnum chryseum, Leskea chrysea, Pylaisia chrysea, Stereodon chryseus]OZB-O03
Orthothecium intricatum Schimp. in Bruch, Schimp. & Gümbel 1851S04 [=Holmgrenia intricataOZB-O03, Isothecium intricatumOZB-O03, Leskea intricataD24, Pylaisia intricataOZB-O03; incl. Orthothecium intricatum var. abbreviatum Dixon 1923S04, D24, Hypnum irroratumOZB-O03, Stereodon subrufusD24]
Orthothecium rufescens (Dicks. ex Brid.) Schimp. in Bruch, Schimp. & Gümbel 1851S04 [=Holmgrenia rufescensOZB-O03, Hypnum rufescensD24, Isothecium rufescensOZB-O03, Leskea rufescensOZB-O03, Pylaisia rufescensOZB-O03, Stereodon rufescensD24]
Plagiothecium sect. Philoscia (Berk.) Ochyra in Ochyra, Żarnowiec & Bednarek-Ochyra 2003 [=Philoscia Berkeley 1863; incl. Plagiotheciella Fleischer in Brotherus 1925 (nom. illeg.)]OZB-O03
Plagiothecium cavifolium (Brid.) Iwats. 1970OZB-O03, S04 [=Hypnum cavifoliumOZB-O03, P. sylvaticum var. cavifoliumOZB-O03; incl. P. roeseanum f. acuminatumOZB-O03, P. roeseanum f. flagellaceumOZB-O03, P. roeseanum f. heterophyllumOZB-O03, P. sylvaticum var. myurumOZB-O03, P. roeseanum f. propaguliferumOZB-O03, Hypnum roeseanumOZB-O03, Plagiothecium lucensOZB-O03, P. roeseanumS04, P. sylvaticum var. roeseiOZB-O03]
Plagiothecium curvifolium Schlieph. ex Limpr. 1897OZB-O03, S04 [=P. denticulatum var. curvifoliumOZB-O03, P. laetum var. curvifoliumM08; incl. P. denticulatum var. aptychusS04, P. denticulatum var. recurvumOZB-O03, P. denticulatum var. secundumOZB-O03]
Plagiothecium denticulatum (Hedw.) Schimp. in Bruch, Schimp. & Gümbel 1851OZB-O03, S04 [=Hypnum denticulatumSK02, Stereodon denticulatusOZB-O03; incl. Plagiothecium denticulatum var. densumOZB-O03, P. denticuatum var. doniiD24, P. donnianumSK02, Stereodon donnianusSK02, P. kinlayanumD24, P. nitellumOZB-O03, P. denticulatum var. orthocladumOZB-O03, P. silvaticum var. orthocladumD24, P. trichodeumD24]
Plagiothecium denticulatum var. undulatum Ruthe ex Geheeb 1877 non P. undulatum (Hedw.) Schimp. in Bruch et al. 1851S04 [incl. P. ruthei Limpr. 1897S04, OZB-O03, P. denticulatum var. rutheiOZB-O03, P. stoloniferumOZB-O03]
Plagiothecium laetum Schimp. in Bruch, Schimp. & Gümbel 1851OZB-O03, S04 [=P. denticulatum var. laetumOZB-O03; incl. P. laetum f. julaceumOZB-O03, P. luridumOZB-O03]
Plagiothecium lamprostachys (Hampe) Jaeger 1878 [=Hypnum lamprostachys; incl. P. novae-seelandiae Broth. 1916]SK02
Plagiothecium latebricola Schimp. in Bruch, Schimp. & Gümbel 1851S04 [=Leskea latebricolaD24, *Philoscia latebricolaOZB-O03, Plagiotheciella latebricolaS04]
Plagiothecium nemorale (Mitt.) Jaeger 1878OZB-O03, S04 [incl. P. neglectumS04, Hypnum sylvaticumOZB-O03, P. sylvaticumOZB-O03, Stereodon sylvaticusOZB-O03]
Plagiothecium piliferum (Schwartz) Schimp. in Bruch et al. 1851S04 [=Isopterygium piliferumS04, Leskea piliferaD24, Plagiotheciella piliferaS04]
Plagiothecium platyphyllum Mönk. 1927OZB-O03, S04 [=P. neglectum ssp. platyphyllumOZB-O03; incl. P. neglectum ssp. platyphyllum f. fontanumOZB-O03]
Plagiothecium succulentum (Wilson) Lindb. 1865OZB-O03, S04 [=Hypnum succulentumOZB-O03, P. denticulatum var. succulentumOZB-O03, P. laetum ssp. succulentumOZB-O03, P. sylvaticum var. succulentumOZB-O03; incl. P. laetum ssp. succulentum f. longifoliumOZB-O03]
Plagiothecium undulatum (Hedw.) Schimp. in Bruch et al. 1851S04 [=Hypnum undulatumD24, *Buckiella undulata (Hedw.) Ireland 2001OZB-O03, Stereodon undulatusOZB-O03; incl. Plagiothecium undulatum f. recurvum non P. denticulatum var. recurvumOZB-O03, P. undulatum f. teresOZB-O03]
*Pseudotaxiphyllum elegans (Brid.) Iwats. 1987OZB-O03, S04 [=Hypnum elegansD24, Isopterygium elegansS04, Isothecium elegansOZB-O03, Plagiothecium elegansS04, Rhynchostegium elegansOZB-O03; incl. Pl. borrerianumD24, Isopterygium elegans var. nanumOZB-O03, Pl. schimperi var. nanumOZB-O03, Isop. elegans var. schimperiOZB-O03, Pl. schimperiOZB-O03, Isop. elegans var. terrestreOZB-O03]
Taxiphyllum densifolium (Lindb. ex Broth.) Reimers 1940 [=Isopterygium densifolium, Plagiothecium densifolium, Rhynchostegium densifolium; incl. Taxiphyllum densifolium var. concavum]OZB-O03
Taxiphyllum taxirameum (Mitt.) Fleischer 1923 [=Isopterygium taxirameum; incl. I. robustum, Taxiphyllum robustum, I. teysmannii]SK02
Taxiphyllum wissgrillii (Garov.) Wijk & Margad. 1960S04 [=Isopterygium wissgrilliiS04; incl. Eurhynchium depressumOZB-O03, Hypnum depressumS04, Isopterygium depressumS04, Plagiothecium depressumS04, Rhynchostegium depressumD24, Taxiphyllum depressumS04]
*Type species of generic name indicated
References
[B-OO10] Bednarek-Ochyra, H., & R. Ochyra. 2010. Bucklandiella shevockii (Bryophyta, Grimmiaceae), an exquisite new species from Yunnan, China. Polish Botanical Journal 55 (2): 499–506.
[BW15] Brotherus, V. F., & W. W. Watts. 1915. The mosses of the New Hebrides [part 2]. Journal and Proceedings of the Royal Society of New South Wales 49 (2): 145–157.
[D24] Dixon, H. N. 1924. The Student’s Handbook of British Mosses 3rd ed. V. V. Sumfield: Eastbourne.
[D03] Dusén, P. 1903. Patagonian and Fuegian mosses. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 63–126. The University: Princeton (New Jersey).
[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.
[M08] Marstaller, R. 2008. Moosgesellschaften am Südrand des Kyffhäusergebirges bei Bad Frankenhausen (Kyffhäuserkreis). 130. Beitrag zur Moosvegetation Thüringens. Mauritiana 20 (2): 289–348.
[N02] Negi, H. R. 2002. Abundance and diversity of moss communities of Chopta-Tunganath in the Garhwal Himalaya. Journal of the Bombay Natural History Society 99 (3): 418–433.
[OZB-O03] Ochyra, R., J. Żarnowiec & H. Bednarek-Ochyra. 2003. Census Catalogue of Polish Mosses. Institute of Botany, Polish Academy of Sciences: Cracow.
Pedersen, N., & L. Hedenäs. 2002. Phylogeny of the Plagiotheciaceae based on molecular and morphological evidence. Bryologist 105 (3): 310–324.
[S04] Smith, A. J. E. 2004. The Moss Flora of Britain and Ireland 2nd ed. Cambridge University Press.
[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.