Belongs within: Polydesmida.
The Platyrhacidae are a family of often large and colourful millipedes found in southeast Asia and Central and South America. Gonopod structure is very simple, with coxae lacking apophyses, prefemora usually lacking processes, and the telopodite usually terminating in a small slender solenomere and larger, shielding tibiotarsal element (Hoffman 2001).
Characters (Hoffman 2001): Ozopores surrounded by broadened discoid peritremata, often at some distance from paranotal margin; epiproct apically broadened; “compound” setae present on labrum (and often also on epiproct). Prozonal texture finely to coarsely granulate. Stigmata, especially the anterior of each pair, distinctly enlarged and elevated. Tibiae of legs with distinctly enlarged, apparently socketed, dorso-apical macroseta. Males with unmodified anterior legs and sterna. Vasa deferentia opening flush on ventral surface of coxae of second pair of legs. Gonopods typically simple in form, without prefemoral process. Gonosternal elements reduced or missing.
<==Platyrhacidae [Euryuridae, Platyrhachidae, Platyrhacoidea, Platyrrhachidae]
|--Platyrhacini [Acanthodesmini]H01
| |--Platyrhacus Hock 1847H01
| | | i. s.: P. amauros Attems 1897H01
| | | P. beaufortiA27
| | |--*P. (Platyrhacus) fuscusH01
| | `--P. (Tirodesmus Cook 1896) [incl. Aymaresmus Chamberlin 1941]K55
| | |--P. (T.) celinusK55
| | |--P. (T.) obscurus Kraus 1955K55
| | |--P. (T.) orellanusK55
| | `--P. (T.) tampocusK55
| |--Stenoniodes Pocock 1897H01
| | |--*S. catorii Pocock 1897H01
| | |--S. angulicollis Pocock 1897 [incl. Platyrhacus bengitanus Attems 1932]H01
| | |--S. arrogans (Attems 1932) [=Platyrhacus arrogans]H01
| | |--S. baluensis Pocock 1897H01
| | |--S. creaghii Pocock 1897H01
| | `--S. sibutensis Pocock 1897H01
| |--Acanthodesmus Peters 1864H01
| | |--*A. pilipes (Peters 1864) [=Polydesmus (*Acanthodesmus) pilipes]H01
| | |--A. anambasius (Attems 1932) [=Platyrhacus anambasius]H01
| | |--A. attenuatior (Chamberlin 1921) [=Platyrrhacus attenuatior]H01
| | |--A. discrepans (Pocock 1897) [=Eurydirorhachis discrepans, Platyrhacus discrepans]H01
| | |--A. dorsalis (Silvestri 1896) [=Platyrhacus dorsalis, P. (Pleorhacus) anthropophagorum Attems 1914]H01
| | |--A. mortoni (Carl 1909) [=Platyrrhacus mortoni]H01
| | `--A. picteti (Silvestri 1899) [=Eurydirorhachis picteti]H01
| |--Eurydirorhachis Pocock 1897H01
| | |--*E. dulitensis Pocock 1897H01
| | |--E. baramanensis Pocock 1897H01
| | `--E. plakodonota (Attems 1897) [=Platyrrhacus plakodonotus]H01
| |--Arbelorhacus Hoffman 2001H01
| | |--*A. magirus Hoffman 2001H01
| | `--A. baramanus (Attems 1897) [=Platyrrhacus baramanus, Platyrhacus baramanus]H01
| |--Barydesmus Cook 1896H01
| |--Nyssodesmus Cook 1896H01
| `--Rhypodesmus Cook 1896H01
| `--R. druryi (Newport 1844) [incl. R. amphelictus Chamberlin 1921, R. terminalis Cook 1897]H01
|--Phyodesmus Cook 1896 [Phyodesmini]H01
| |--*P. pictus (Peters 1864) [=Polydesmus pictus]H01
| |--P. hosei Pocock 1897H01
| |--P. laticlavius Hoffman 1978H01
| |--P. longispinosus (Silvestri 1896)H01
| |--P. magnificus (Silvestri 1896) [=Platyrhacus magnificus]H01
| |--P. ornatus Pocock 1897H01
| |--P. scutatus (Peters 1864)H01
| `--P. sublimbatus (Silvestri 1897)H01
|--HoplurorhachiniH01
| |--Hoplurorhachis Pocock 1897H01
| | |--*H. everettii Pocock 1897 [=Platyrhacus (Psaphodesmus) everettii, Platyrrhacus (Pleorhacus) everettii]H01
| | |--H. crassipes (Carl 1909) (see below for synonymy)H01
| | `--H. oeceter Hoffman 2001H01
| |--Creagronopus Hoffman 2001H01
| | |--*C. hosei (Pocock 1897) (see below for synonymy)H01
| | `--C. lamprus (Chamberlin 1921) [=Platyrrhacus lamprus, Platyrhacus (Psaphodesmus) lamprus]H01
| `--Sabarhacus Hoffman 2001H01
| |--*S. derodontus Hoffman 2001H01
| |--S. gongylodes (Attems 1899) [=Platyrrhacus gongylodes]H01
| `--S. sucidus (Attems 1932) [=Platyrhacus sucidus, P. (Psaphodesmus) sucidus]H01
`--PsaphodesminiH01
|--Parazodesmus verrucosusH01
|--ZodesmusH01
`--OzorhacusH01
Platyrhacidae incertae sedis:
EuryurusH90 [EuryurinaeH01]
|--E. leachiiS96
`--E. maculatusH90
Auturus erythropygosH90, S96 [=Euryurus erythropygusS96]
Amplinus Attems 1901K54
|--A. klugiiK54
|--A. palicaudatus (Attems 1901) [=Pachyurus palicaudatus]K54
`--A. serratus Kraus 1954K54
DerodesmusH01
Polylepiscus campanulaeH01
HaplorhacusH01
|--*H. haplopusH01
`--H. doryphorus (Attems 1899) (n. d.) [=Platyrhacus (Haplorhacus) doryphorus]H01
*Creagronopus hosei (Pocock 1897) [=Hoplurorhachis hosei non Phyodesmus hosei Pocock 1897 (not preoc. if in dif. gen.), Platyrhacus (Psaphodesmus) hosei, Platyrrhacus hoplurorhachis Attems 1899, Platyrhacus (Pleorhacus) hoplurorhachis]H01
Hoplurorhachis crassipes (Carl 1909) [=Platyrrhacus crassipes, Platyrhacus (Pleorhacus) crassipes, Platyrhacus (Psaphodesmus) crassipes]H01
*Type species of generic name indicated
References
[A27] Attems, C. 1927. Myriopoden von Ambon und anderen Inseln der Banda-See. Zoologische Mededeelingen 10: 61–70.
[H90] Hoffman, R. L. 1990. Diplopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 835–860. John Wiley & Sones: New York.
[H01] Hoffman, R. L. 2001. A synopsis of the platyrhacid millipeds of Borneo (Diplopoda: Polydesmida: Platyrhacidae). Revue Suisse de Zoologie 108 (2): 403–440.
[K54] Kraus, O. 1954. Myriapoden aus El Salvador. Senckenbergiana Biologica 35 (5–6): 293–349.
[K55] Kraus, O. 1955. Myriapoden aus Peru, III. Senckenbergiana Biologica 36 (3–4): 173–200.
[S96] Shelley, R. M. 1996. Book Review: The Biology of Millipedes. Stephen P. Hopkin and Helen J. Read. Proceedings of the Entomological Society of Washington 98 (1): 169–172.